Haakon Hop » Members

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Haakon Hop

Senior research scientist Norwegian Polar InstituteResearch department: ecotoxicology
Work address Norwegian Polar Institute Fram Centre Tromsø 9296 Norway Work Phone: +47 77 75 05 22
Photo of Haakon Hop


Research interests:

  • Ecology and bioenergetics of pelagic, sympagic (ice-associated) and hard-bottom invertebrates and fishes in Arctic marine environments
  • Marine ecosystem structure (stable isotopes), energy flow (lipids), and bioaccumulation of persistent pollutants in Arctic marine food webs
  • Integration of physical and biological data in multidisciplinary research projects carried out in the Marginal Ice Zone (MIZ) and fjords in Svalbard

My experience includes >25 years of Arctic research, both experimental and field based work. I have organized multidisciplinary research programs and cruises, and participated on these as researcher, scientific diver and cruise leader. I have published 130 papers in the peer-reviewed literature, including many multidisciplinary papers. I also have extensive teaching experience as course instructor, lecturer and external examiner at the university level, and am external supervisor for graduate students. I have organized and led workshops and sat on review panels for international research proposals. I have lead international groups for establishing new marine research projects and a new Arctic Marine Laboratory in Ny-Ålesund. I am currently leading marine ecological studies in ICE – Centre for ice, climate and environment.

Running national and international projects:

  • MOSJ-ICE-project to determine inter-annual variation in hydrography and pelagic plankton in Kongsfjorden and Rijpfjorden in Svalbard (NPI)
  • N-ICE2015 participation in the Arctic Ocean with dive team to determine under-ice flora and fauna (NPI)
  • Ice-algal and under-ice phytoplankton bloom dynamics in a changing Arctic icescape (Norwegian Research Council)

Completed national and international projects:

  • Kongsfjorden Ecosystem – new views after more than a decade of research (NRC)
  • Impact of ocean acidification on Arctic zooplankton populations (NRC)
  • Response of marine and terrestrial ecosystems to climate changes in Arctic – links between physical environment biodiversity of zooplankton and seabird populations (ALKEKONGE) (Polish-Norwegian Research Fund)
  • Marine ecosystem consequences of climate induced changes in water masses off West-Spitsbergen (MariClim) (NRC)

Relevant scientific publications:

  1. Hop, H. and C. Wiencke. 2019. Editorial: The ecosystem of Kongsfjorden, Svalbard. Advances in Polar Ecology (accepted).
  2. Hop, H., F. Cottier, and J. Berge. 2019. Autonomous marine observatories in Kongsfjorden, Svalbard. Advances in Polar Ecology (accepted).
  3. Hop, H., A. Wold, M. Vihtakari, M. Daase, S. Kwasniewski, M. Gluchowska, S. Lischka, F. Buchholz, and S. Falk-Petersen. 2019. Zooplankton in Kongsfjorden (1996-2016) in relation to climate change. Advances in Polar Ecology (accepted).
  4. Pavlova, O, S. Gerland, and H. Hop. 2019. Changes in sea-ice extent and thickness in Kongsfjorden, Svalbard (2003-2016). Advances in Polar Ecology.
  5. Bischof, K., P. Convey, P. Duarte, J.-P. Gattuso, M. Granberg, H. Hop, C. Hoppe, C. Jiménez, L. Lisitsyn, B, Martínez, M.Y. Roleda, P. Thor, J. Wiktor, and G.W. Gabrielsen. 2019. Kongsfjorden as harbinger of the future Arctic: knowns, unknowns and research priorities. Advances in Polar Ecology (accepted).
  6. Duarte, P., J.M. Weslawski, and H. Hop. 2019. Outline of an Arctic fjord ecosystem model for Kongsfjorden-Krossfjorden, Svalbard. Advances in Polar Ecology (accepted).
  7. Fredriksen, S., U. Karsten, I. Bartsch, J. Woelfel, M. Koblowsky, R. Schumann, S.R. Moy, R.S. Steneck, J. Wiktor, H. Hop, C. Wiencke. 2019. Biodiversity of benthic macro- and microalgae from Svalbard with special focus on Kongsfjorden. Advances in Polar Ecology (accepted).
  8. Rastrick, S.S.P, H.E. Graham, K. Azetsu-Scott, P. Calosi, M. Chierici, A. Fransson, H. Hop, J. Hall-Spencer, M. Milazzo, P. Thor, and T. Kutti. 2018. Using natural analogues to investigate the effects of climate change and ocean acidification on Northern ecosystems. ICES Journal of Marine Science 75: 2299-2311.
  9. Fernández-Méndez, M., L.M. Olsen, H.M. Kauko, A. Meyer, A. Rösel, I. Merkouriadi, C.J. Mundy, J.K. Ehn, A.M. Johansson, P.M. Wagner, Å. Ervik, B.K. Sorrell, P.M. Duarte, A. Wold, H. Hop, and P. Assmy. 2018. Algal hot spots in a changing Arctic Ocean: Sea-ice ridges and the snow-ice interface. Frontiers Marine Science 5:75. Doi: 10.3389/fmars.2018.00075.
  10. Meyer Ottesen. C.A., H. Hop, J.S. Christiansen, and Stig Falk-Petersen. 2018. Reproduction and sexual dimorphism of daubed shanny (Teleostei: Leptoclinus maculatus) in Svalbard waters. Polar Biology 41: 1867-1880.
  11. Vihtakari, M., J. Welcker, B. Moe, O. Chastel, S. Tartu, H. Hop, C. Bech, S. Descamps, and G.W. Gabrielsen. 2018. Black-legged kittiwakes as messengers of Atlantification in the Arctic. Scientific Reports 8, 1178. Doi: 0.1038/s41598-017-19118-8.
  12. Bluhm, B.A., H. Hop, M. Vihtakari, R. Gradinger, K. Iken, I.A. Melnikov, and J.E. Søreide. 2018. Sea ice meiofauna distribution on local to pan-Arctic scales. Ecology and Evolution 8: 2350-2364.
  13. Olsen, L.M., S.R. Laney, P. Duarte, H.M. Kauko, M. Fernández-Méndez, C.J. Mundy, A. Rösel, A. Meyer, P. Itkin, L. Cohen, I. Peeken, A. Tatarek, M. Rozanska, J. Wiktor, T. Taskjelle, A.K. Pavlov, S.R. Hudson, M.A. Granskog, H. Hop, P. Assmy. 2017. The seeding of ice-algal blooms in Arctic pack ice: the multiyear ice seed repository hypothesis. Journal of Geophysical Research-Biogeosciences 122: 1529-1548.
  14. Duarte, P., A. Meyer, L.M. Olsen, H.M. Kauko, P. Assmy, A. Rösel, P. Itkin, S.R. Hudson, M.A. Granskog, S. Gerland, A. Sundfjord, H. Steen, H. Hop, N. Jeffery, S.M. Elliott, E.C. Hunke, and A.K. Turner. 2017. Sea-ice thermohaline-dynamics and biogeochemistry in the Arctic Ocean: empirical and model results. Journal of Geophysical Research-Biogeosciences 122: 1632-1654.
  15. Brown, T.A., P. Assmy, H. Hop, A. Wold, and S.T. Belt. 2017. Transfer of ice algae carbon to ice-associated amphipods in the high-Arctic pack ice environment. Journal of Plankton Research 39: 664-674.
  16. Bailey, A., P. De Wit, P. Thor, H.I. Browman, R. Bjelland, S. Shema, D.M. Fields, J.A. Runge, C. Thompson, and H. Hop. 2017. Regulation of gene expression is associated with tolerance of the Arctic copepod Calanus glacialis to CO2-acidified sea water. Ecology and Evolution 7: 7145-1760.
  17. Haug, T., S. Falk-Petersen, M. Greenacre, H. Hop, U. Lindstrøm, S. Meier, K.T. Nilssen, and A. Wold. 2017. Trophic levels and fatty acids i
  18. Assmy, P., M. Fernández-Méndez, P. Duarte, A. Meyer, A. Randelhoff, C.J. Mundy, L.M. Olsen, H.M. Kauko, A. Bailey, M. Chierici, L. Cohen, A.P. Doulgeris, J.K. Ehn, A. Fransson, S. Gerland, H. Hop, S.R. Hudson, N. Hughes, P. Itkin, G. Johnsen, J.A. King, B.P. Koch, Z. Koenig, S. Kwasniewski, S.R. Laney, M. Nicolaus, A.K. Pavlov, C.M. Polashenski, C. Provost, A. Rösel, M. Sandbu, G. Spreen, L.H. Smedsrud, A. Sundfjord, T. Taskjelle, A. Tatarek, J. Wiktor, P.M. Wagner, A. Wold, H. Steen, and M.A. Granskog. 2017. Leads in Arctic pack ice enable early phytoplankton blooms below snow-covered sea ice. Scientific Reports 7, 40850; doi: 10.1038/srep40850.
  19. Duarte, P., J.M. Weslawski, and H. Hop. 2017. Outline of an Arctic fjord ecosystem model for Kongsfjorden-Krossfjorden, Svalbard. Advances in Polar Ecology (accepted).
  20. Fredriksen, S., U. Karsten, I. Bartsch, J. Woelfel, M. Koblowsky, R. Schumann, S.R. Moy, R.S. Steneck, J. Wiktor, H. Hop, C. Wiencke. 2017. Biodiversity of benthic macro- and microalgae from Svalbard with special focus on Kongsfjorden. Advances in Polar Ecology (accepted).
  21. Bailey, A., P. Thor, H.I. Browman, D.M. Fields, J. Runge, A. Vermont, R. Bjelland, C. Thompson, S. Shema, C.M.F. Durif, and H. Hop. 2017. Early life stages of the Arctic copepod Calanus glacialis are unaffected by increased seawater pCO2. ICES Journal of Marine Science. Doi: 10.1093/icesjms/fsw066
  22. Vihtakari, M., W.G. Ambrose Jr., P.E. Renaud, W.L. Locke V, M.L. Carroll, J. Berge, L.J. Clarke, F. Cottier, and H. Hop. 2016. A key to the past? Element ratios as environmental proxies in two Arctic bivalves. Palaeogeography, Palaeoclimatology, Palaeoecology 465: 316-332.
  23. Hunt Jr., G.L., K.F. Drinkwater, K. Arrigo, J. Berge, K.L. Daly, S. Danielson, M. Daase, H. Hop, E. Isla, N. Karnovsky, K. Laidre, E.J. Murphy, F.J. Mueter, P.E. Renaud, W.O. Smith Jr., P. Trathan, J. Turner, and D. Wolf-Gladrow. 2016. Advection in polar and sub-polar environments: Impacts on high latitude marine ecosystems. Progress in Oceanography 149: 40-81.
  24. Wiencke, C. and H. Hop. 2016. Editorial: Ecosystem Kongsfjorden– new views after more than a decade of research. Polar Biology 39: 1679-1687.
  25. Gluchowska M., S. Kwasniewski, A. Prominska, A. Olszewska, I. Goszczko, S. Falk-Petersen, H. Hop, and J.M. Weslawski. 2016. Zooplankton in Svalbard fjords on the Atlantic-Arctic boundary. Polar Biology 39: 1785-1802.
  26. Fransson A., M. Chierici, H. Hop, H.S. Findlay, S. Kristiansen, and A. Wold. 2016. Late winter-to-summer change in ocean acidification state in Kongsfjorden, with implications for calcifying organisms. Polar Biology 39: 1841-1857.
  27. Voronkov, A., H. Hop, and B. Gulliksen. 2016. Zoobenthic communities on hard-bottom habitats in Kongsfjorden, Svalbard. Polar Biology 39: 2077-2095.
  28. Vihtakari, M., P.E. Renaud, L.J. Clarke, M.J. Whitehouse, H. Hop, M.L. Carroll, and W.G. Ambrose Jr. 2016. Decoding the oxygen isotope signal for seasonal growth patterns in Arctic bivalves. Palaeogeography, Palaeoclimatology, Palaeoecology 446:263-283.
  29. Dalpadado, P., H. Hop, J. Rønning, V. Pavlov, E. Sperfeld, F. Buchholz, A. Rey, and A. Wold. 2016 Distribution and abundance of euphausiids and pelagic amphipods in Kongsfjorden, Isfjorden and Rijpfjorden (Svalbard) and changes in their importance as key prey in a warming marine ecosystem. Polar Biology. Kongsfjorden Special Issue (accepted)
  30. Bartsch I, M. Paar, S. Fredriksen, M. Schwanitz, C. Daniel, H. Hop, and C. Wiencke. 2016. Changes in kelp forest biomass and depth distribution in Kongsfjorden, Svalbard, between 1996-1998 and 2012-2014 reflect Arctic warming. Polar Biology, Kongsfjorden Special Issue (accepted).
  31. Daase, M., H. Hop, and S. Falk-Petersen. 2016. Small-scale diel vertical migration of zooplankton in the High Arctic. Polar Biology. doi:10.1007/s00300-015-1840-7.
  32. Fredriksen, S., U. Karsten, I. Bartsch, J. Woelfel, M. Koblowsky, R. Schumann, S.R. Moy, R.S. Steneck, J. Wiktor, Hop, C. Wiencke. 2016. Biodiversity of benthic macro- and microalgae from Svalbard with special focus on Kongsfjorden. Advances in Polar Ecology (accepted).
  33. Paar, M, A. Voronkov, Hop, T. Brey, I. Bartsch, M. Schwanitz, C. Wiencke, B. Lebreton, R. Asmus, and H. Asmus. 2016. Temporal shift in biomass and production of macrozoobenthos in the macroalgal belt at Hansneset, Kongsfjorden, after 15 years. Polar Biology. doi: 10.1007/s00300-015-1760-6.
  34. Barber, D.G., Hop, C.J. Mundy, B. Else, I.A. Dmitrenko, J.-E. Tremblay, J.K. Ehn, P. Assmy, M. Daase, L.M. Candlish, and S. Rysgaard. 2015. Selected physical, biological and biogeochemical implications of a rapidly changing Arctic Marginal Ice Zone. Progress in Oceanography 139:122-150.
  35. Duarte, P., P. Assmy, Hop, G. Spreen, S. Gerland, and S.R. Hudson. 2015. The importance of vertical resolution in sea ice algae production models. Journal of Marine Systems 145: 69-90.
  36. Berge, J., K. Heggland, O.J. Lønne, F. Cottier, Hop, G.W. Gabrielsen, L. Nøttestad, and O.A. Misund. 2015. First records of Atlantic mackerel (Scomber scombrus) from the Svalbard archipelago, Norway, with possible explanations for the extension of its distribution. Arctic 68: 54-61.
  37. Lynghammar, A., J.S. Christiansen, A.M. Griffiths, S.-E. Fevolden, Hop, and T. Bakken. 2014. DNA barcoding of the northern Northeast Atlantic skates (Chondrichthyes, Rajidae), with remarks on the widely distributed starry ray. Zoologica Scripta 43: 485-495.
  38. Meyer Ottesen, C.A., Hop, S. Falk-Petersen, and J.S. Christiansen. 2014. Growth of daubed shanny (Teleostei: Leptoclinus maculatus) in Svalbard waters. Polar Biology 37: 809-815.
  39. Hovinen, J.E.H., K. Wojczulanis-Jakubas, D. Jakubas, Hop, J. Berge, D. Kidawa, N..J. Karnovsky, and H. Steen. 2014. Fledging success of a little auks in the high Arctic: do provisioning rates and the quality of foraging grounds matter?  Polar Biology 37: 665-674.
  40. Hovinen, J.E.H., J. Welcker, A. Rabindranath, W. Brown, H. Hop, J. Berge, and H. Steen. 2014. At-sea distribution of foraging little auks relative to physical factors and potential food supply. Marine Ecology Progress Series 503: 263-277.
  41. Murzina, S.A., Z.A. Nefedova, S. Falk-Petersen, Hop, T.R. Ryokolainen, C.A. Meyer Ottesen, P.O. Ripatti, J. Berge, and N.N. Nemova. 2013. Lipids in the daubed shanny (Teleostei: Leptoclinus maculatus) in Svalbard. Polar Biology 36: 1619-1631.
  42. Assmy, P., J.K. Ehn, M. Fernández-Méndez, Hop, C. Katlein, A. Sundfjord, K. Bluhm, M. Daase, A. Engel, , A. Fransson,  M.A. Granskog, S.R. Hudson, S. Kristiansen, M. Nicolaus, I. Peeken, A.H.H. Renner, G. Spreen, A. Tatarek, and  J. Wiktor. 2013. Floating ice-algal aggregates below melting Arctic sea ice. PloS ONE 8 (10), e76599.
  43. McMeans, B.C., M.T. Arts, C. Lydersen, K.M Kovacs, Hop, S. Falk-Petersen, and A.T. Fisk. 2013. The role of Greenland sharks (Somniosus microcephalus) in an arctic ecosystem – assessed via stable isotopes and fatty acids. Marine Biology 160: 1223-1238.
  44. Søreide, J.E., M.L. Carroll, Hop, W.G. Ambrose Jr., E.N. Hegseth, and S. Falk-Petersen. 2013. Sympagic-pelagic-benthic coupling in Arctic and Atlantic waters around Svalbard revealed by stable isotopic and fatty acid tracers. Marine Biology Research 9: 831-850.
  45. Hop, H. and H. Gjøsæter. 2013. Polar cod (Boreogadus saida) and capelin (Mallotus villosus) as key species in marine food webs of the Arctic and the Barents Sea. Marine Biology Research 9: 878-894.
  46. Voronkov, A, Hop, and B. Gulliksen. 2013. Diversity of hard-bottom fauna relative to environmental gradients in Kongsfjorden, Svalbard. Polar Research 32, doi :10.30402/polar.v32i0.11208. Supplementary Table S1.
  47. Hindell, M.A., C. Lydersen, Hop, and K. M. Kovacs. 2012. Pre-partum diet of adult female bearded seals in years of contrasting ice conditions. PLoS ONE 7 (5):1-10.
  48. Hop, H., C. Wiencke, B. Võgele, and N.A. Kovaltchouk. 2012. Species composition, zonation and biomass of marine benthic macroalgae in Kongsfjorden, Svalbard. Botanica Marina 55: 399-414.
  49. Christiansen, J.S., Hop, E.M. Nilssen, and J. Joensen. 2012. Trophic ecology of sympatric Arctic gadoids, Arctogadus glacialis (Peters, 1872) and Boreogadus saida (Lepechin, 1774), in NE Greenland. Polar Biology 35:1247-1257.
  50. Trudnowska, E., J. Szczucka, L. Hoppe, R. Boehnke, Hop, and K. Blachowiak- Samolyk. 2012. Multidimensional zooplankton observations on the northern West Spitsbergen Shelf. Journal of Marine Systems 98-99: 18-25.
  51. Murzina, S.A., C.A. Meyer Ottesen, S. Falk-Petersen, Hop, N.N. Nemova, and O.G. Poluektova. 2012. Oogenesis and lipids in gonad and liver of daubed shanny (Leptoclinus maculatus) females from Svalbard waters. Fish Physiology and Biochemistry 38:1393-1407.
  52. Vieweg, I, Hop, T. Brey, S. Huber, W. G. Ambrose Jr., W.L. Locke V, and G.W. Gabrielsen. 2012. Persistent organic pollutants in four bivalve species from Svalbard waters. Environmental Pollution 161: 134-142.
  53. Hodal, H., S. Falk-Petersen, Hop, S. Kristiansen and M. Reigstad. 2012. Spring bloom dynamics in Kongsfjorden, Svalbard: Nutrients, phytoplankton, protozoans and primary production. Polar Biology 35: 191-203.
  54. Hop, H., Haug, and S. Falk-Petersen. 2011. Editorial: Arctic Marine Biodiversity under Change. Marine Biodiversity 41: 349-351.
  55. Ehn, J.K., C.J. Mundy, D.G. Barber, Hop, A. Rossnagel, and J. Stewart. 2011. Impact of horizontal spreading on light propagation in melt pond covered seasonal sea ice in the Canadian Arctic. Journal of Geophysical Research. 116, C00G02, doi:10.1029/2010JC006908.
  56. Carroll, M.L., W.G. Ambrose Jr., B.S. Levin, W.L. Locke V, G.A. Henkes, Hop, and P.E. Renaud. 2011. Pan-Svalbard growth rate variability and environmental regulation in the Arctic bivalve Serripes groenlandicus. Journal of Marine Systems 88: 239-251.
  57. Ki, J-S., H.-U. Dahms, I.-C. Kim, H.G. Park, Hop, and J.-S. Lee. 2011. Molecular relationships of gammaridean amphipods from Arctic sea ice. Polar Biology 34: 1559-1569.
  58. Mundy, C.J., M. Gosselin, J.K. Ehn, C. Belzile, M. Poulin, E. Alou, S. Roy, Hop, S. Lessard, T.N. Papakyriakou, D.G. Barber, and J. Stewart. 2011. Characteristics of two distinct high-light acclimated microbial communities during advanced stages of sea ice melt. Polar Biology 34: 1869-1886.
  59. Hop, H., C.J. Mundy, M. Gosselin, A.L. Rossnagel, and D.G. Barber. 2011. Zooplankton boom and ice amphipod bust below melting sea ice in the Amundsen Gulf, Arctic Canada. Polar Biology 34: 1947-1958.
  60. Meyer Ottesen, C.A., Hop, J.S. Christiansen, and S. Falk-Petersen. 2011. Early life history of daubed shanny (Teleostei: Leptoclinus maculatus) in Svalbard waters. Marine Biodiversity 41: 383-394.
  61. Wold, A., I. Jæger, Hop, G.W. Gabrielsen, and S. Falk-Petersen. 2011. Arctic seabird food chains explored by fatty acid composition and stable isotopes in Kongsfjorden, Svalbard. Polar Biology 34:1147-1155.
  62. Petursdottir, H., S. Falk-Petersen, H. Hop, and A. Gislason. 2010. Calanus finmarchicus along the northern Mid-Atlantic Ridge: variation in fatty acid and alcohol profiles and stable isotopes values, δ15N and δ13 Journal of Plankton Research 32: 1067-1077.
  63. Nahrgang, J., L. Camus, F. Broms, J.S. Christiansen, and Hop. 2010. Seasonal baseline levels of physiological and biochemical parameters in polar cod (Boreogadus saida): Implication for environmental monitoring. Marine Pollution Bulletin 60: 1336-1345.
  64. Ki, J.-S., Hop, S-J. Kim, I.-C. Kim, H.G. Park, and J.-S. Lee. 2010. Complete mitochondrial genome in the Arctic gammaridean, Onisimus nanseni (Crustacea; Amphipoda): novel gene structures and unusual control region features. Comparative Biochemistry and Physiology, Part D, 5: 105-115.
  65. Voronkov, A., S.D. Stepanjants, and Hop. 2010. Hydrozoan diversity on hard bottom in Kongsfjorden. Journal of the Marine Biological Association of the United Kingdom 90: 69-84.
  66. Nahrgang, J., L. Camus, P. Gonzalez, M. Jönsson, J.S. Christiansen, and Hop. 2010. Biomarker responses in polar cod (Boreogadus saida) exposed to dietary crude oil. Aquatic Toxicology 96: 77-83.
  67. Nahrgang, J., L. Camus, M.G. Carls, P. Gonzalez, M. Jönsson, I.C. Taban, R.K. Bechmann, J.S. Christiansen, and Hop. 2010. Biomarker responses in polar cod (Boreogadus saida) exposed to the water soluble fraction of crude oil. Aquatic Toxicology 97: 234-242.
  68. Nahrgang, J., L. Camus, P. Gonzalez, A. Goksøyr, J.S. Christiansen, and Hop. 2009. PAH biomarker responses in polar cod (Boreogadus saida) exposed to benzo(a)pyrene. Aquatic Toxicology 94: 309-319.
  69. Jæger, I., Hop, and G.W. Gabrielsen. 2009. Biomagnification of mercury in selected species from an Arctic marine food web in Svalbard. Science of the Total Environment 407: 4744-4751.
  70. Tamelander, T., M. Reigstad, Hop, and T. Ratkova. 2009. Ice algal assemblages and vertical export of organic matter from sea ice in the Barents Sea and Nansen Basin (Arctic Ocean). Polar Biology 32: 1261-1273.
  71. Walkusz, W., S. Kwasniewski, S. Falk-Petersen, Hop, V. Tverberg, P. Wieczorek, and J. M. Weslawski. 2009. Seasonal and spatial changes in the zooplankton community in Kongsfjorden, Svalbard. Polar Research 28: 254-281.
  72. Piwosz, K., W. Walkusz, R. Hapter, P. Wieczorek, Hop, and J. Wiktor. 2009. Comparison of productivity and phytoplankton in warm (Kongsfjorden) and a cold (Hornsund) Spitsbergen fjord in mid-summer 2002. Polar Biology 32: 549-559.
  73. Falk-Petersen, S., T. Haug, Hop, K.T. Nilssen, and A. Wold. 2009. Transfer of lipids from plankton to blubber of harp and hooded seals off East Greenland. Deep-Sea Research II 56: 2080-2086.
  74. Blachowiak-Samolyk, K., J.E. Søreide, S. Kwasniewski, A. Sundfjord, Hop, S. Falk-Petersen, and E. N. Hegseth. 2008. Hydrodynamic control of mesozooplankton abundance and biomass in northern Svalbard waters (79-81°N). Deep-Sea Research II 55: 2210-2224.
  75. Søreide, J.E., S. Falk-Petersen, E.N. Hegseth, Hop, M.L. Carroll, K.A. Hobson, and K. Blachowiak-Samolyk. 2008. Seasonal feeding strategies of Calanus in the high-Arctic Svalbard region. Deep-Sea Research II 55: 2225-2244.
  76. Hop, H. and O. Pavlova. 2008. Distribution and biomass transport of ice amphipods in drifting sea ice around Svalbard. Deep-Sea Research II 55: 2292-2307.
  77. Tamelander, T., M. Reigstad, Hop, M.L. Carroll, and P. Wassmann. 2008. Pelagic and sympagic contribution of organic matter to zooplankton and vertical export in the Barents Sea marginal ice zone. Deep-Sea Research II 55: 2330-2339.
  78. Blachowiak-Samolyk, K., S. Kwasniewski, Hop, and S. Falk-Petersen. 2008. Magnitude of mesozooplankton variability: A case study from the Marginal Ice Zone of the Barents Sea in spring. Journal of Plankton Research 30: 311-323.
  79. Petursdottir, H., A. Gislason, S. Falk-Petersen, Hop, and J. Svavarsson. 2008. Trophic interactions of the pelagic ecosystem over the Reykjanes Ridge as evaluated by fatty acid and stable isotope analyses. Deep-Sea Research II 55: 83-93.
  80. Graeve, M., M. Lundberg, M. Böer, G. Kattner, Hop, and S. Falk-Petersen. 2008. The fate of dietary lipids in the Arctic ctenophore Mertensia ovum (Fabricius 1780). Marine Biology 153: 643-651.
  81. Blachowiak-Samolyk, K., S. Kwasniewski, K. Dmoch, Hop, and S. Falk-Petersen. 2007. Trophic structure of zooplankton in the Fram Strait in spring and autumn 2003. Deep-Sea Research II, 2716-2728.
  82. Fuglei, E., J.O. Bustnes, Hop, T. Mørk, H. Björnfoth, and B. van Bavel. 2007. Environmental contaminants in arctic foxes (Alopex lagopus) in Svalbard: Relationships with feeding ecology and body condition. Environmental Pollution 146: 128-138.
  83. Haukås, M., U. Berger, Hop, B. Gulliksen, and G.W. Gabrielsen. 2007. Bioaccumulation of per- and polyfluorinated alkyl substances (PFAS) in selected marine species from the Barents Sea food web. Environmental Pollution 148: 360-371.
  84. Borgå, K., Hop, J.U. Skaare, H. Wolkers, and G.W. Gabrielsen. 2007. Selective bioaccumulation of chlorinated pesticides and metabolites in Arctic seabirds. Environmental Pollution 145: 545-553.
  85. Søreide, J.E., T. Tamelander, Hop, K.A. Hobson, and I. Johansen. 2006. Sample preparation effects on stable C and N isotope values: a comparison of methods in Arctic marine food web studies. Marine Ecology Progress Series 328: 17-28.
  86. Wassmann, P., M. Reigstad, T. Haug, B. Rudels, M.L. Carroll, Hop, G.W. Gabrielsen, S. Falk-Petersen, S.G. Denisenko, E. Arashkevich, D. Slagstad, and O. Pavlova. 2006. Food webs and carbon flux in the Barents Sea. Progress in Oceanography 71: 232-287.
  87. Mueller, R.P., R.S. Brown, Hop, and L. Moulton. 2006. Video and acoustic camera techniques for studying fish under ice: a review and comparison. Reviews in Fish Biology and Fisheries 16: 213-226.
  88. Hop, H., S. Falk-Petersen, H. Svendsen, S. Kwasniewski, V. Pavlov, O. Pavlova, and J.E. Søreide. 2006. Physical and biological characteristics of the pelagic system across Fram Strait to Kongsfjorden. Progress in Oceanography 71: 182-231.
  89. Søreide, J.E., Hop, M.L. Carroll, S. Falk-Petersen, and E.N. Hegseth. 2006. Seasonal food web structures and sympagic-pelagic coupling in the European Arctic revealed by stable isotopes and a two-source food web model. Progress in Oceanography 71: 59-87.  Søreide et al. 2007, Corrigendum. Progress in Oceanography 73: 96-98.
  90. Dale, K., S. Falk-Petersen, Hop, and S.-E. Fevolden. 2006. Population dynamics and body composition of the Arctic hyperiid amphipod Themisto libellula in Svalbard fjords. Polar Biology 29: 1063-1070.
  91. Lundberg, M., Hop, K. Eiane, B. Gulliksen, and S. Falk-Petersen. 2006. Population structure and accumulation of lipids in the ctenophore Mertensia ovum. Marine Biology 149: 1345-1353.
  92. Tamelander, T., J.E. Søreide, Hop, and M.L. Carroll. 2006. Fractionation of stable isotopes in the Arctic marine copepod Calanus glacialis: Effects on the isotopic composition of marine particulate organic matter. Journal of Experimental Marine Biology and Ecology 333: 231-240.
  93. Sørmo, E.G., M.P. Salmer, B.M. Jenssen, Hop, K. Bæk, K.M. Kovacs, C. Lydersen, S. Falk-Pettersen, G.W. Gabrielsen, E. Lie, and J.U. Skaare. 2006. Biomagnification of polybrominated diphenyl ether and hexabromocyclododecane flame retardants in the polar bear food chain in Svalbard, Norway. Environmental Toxicology and Chemistry 25: 2502-2511.
  94. Tamelander, T., P.E. Renaud, Hop, M.L. Carroll, W.G. Ambrose Jr., and K.A. Hobson. 2006. Trophic relationships and pelagic-benthic coupling during summer in the Barents Sea Marginal Ice Zone, revealed by stable carbon and nitrogen isotope measurements. Marine Ecology Progress Series 310: 33-46.
  95. Blachowiak-Samolyk, K., S. Kwasniewski, K. Richardson, K. Dmoch, E. Hansen, Hop, S. Falk-Petersen, and L.T. Mouritsen. 2006. Arctic zooplankton do not perform diel vertical migration (DVM) during periods of midnight sun. Marine Ecology Progress Series 308: 101-116.
  96. Böer, M., C. Gannefors, G. Kattner, M. Graeve, Hop, and S. Falk-Petersen. 2005. The Arctic pteropod Clione limacina: seasonal lipid dynamics and life strategy. Marine Biology 147: 707-717.
  97. Gannefors, C. M. Böer, G. Kattner, M. Graeve, K. Eiane, B. Gulliksen, Hop, and S. Falk-Petersen. 2005. The Arctic sea butterfly Limacina helicina; lipids and life strategy. Marine Biology 147: 169-177.
  98. Borgå, K., H. Wolkers, J.U. Skaare, Hop, D.C.G. Muir, and G.W. Gabrielsen. 2005. Bioaccumulation of PCBs in Arctic seabirds; influence of dietary exposure and congener biotransformation. Environmental Pollution 134: 397-409.
  99. Walkusz, W., S. Kwasniewski, K. Dmoch, M. I. Zmijewska, S. Falk-Petersen, Hop, and J. Sicinski. 2004. Characteristics of the Arctic and Antarctic mesozooplankton in the neritic zone during summer. Polish Polar Research 25: 275-291.
  100. Wiencke, C., B. Vögele, N.A. Kovaltchouk, and Hop. 2004. Species composition and zonation of marine benthic macroalgae at Hansneset in Kongsfjorden, Svalbard. Pp. 55-62 In: C. Wiencke (ed.). The coastal ecosystem of Kongsfjorden, Svalbard. Synopsis of biological research performed at the Koldewey-Station in the years 1991-2003. Reports on Polar and Marine Research 492.
  101. Engelsen, O., Hop, E.N. Hegseth, E. Hansen, and S. Falk-Petersen. 2004. Deriving phytoplankton biomass in the Marginal Ice Zone from satellite observable parameters. International Journal of Remote Sensing 25: 1453-1457.
  102. Søreide, J.E., Hop, S. Falk-Petersen, B. Gulliksen, and E. Hansen. 2003. Macrozooplankton communities and environmental variables in the Barents Sea marginal ice zone in late winter and spring. Marine Ecology Progress Series 263: 43-64.
  103. Dahl, T.M., S. Falk-Petersen, G.W. Gabrielsen, J.R. Sargent, Hop, and R.-M. Millar. 2003. Lipids and stable isotopes in common eider, black-legged kittiwake and northern fulmar: a trophic study from an Arctic fjord. Marine Ecology Progress Series 256: 257-269.
  104. Kwasniewski, S., Hop, S. Falk-Petersen, and G. Pedersen. 2003. Distribution of Calanus species in Kongsfjorden, a glacial fjord in Svalbard. Journal of Plankton Research 25: 1-20.
  105. Hop, H., T. Pearson, E.N. Hegseth, K.M. Kovacs, C. Wiencke, S. Kwasniewski, K. Eiane, F. Mehlum, B. Gulliksen, M. Wlodarska-Kowalczuk, C. Lydersen, J.M. Weslawski, S. Cochrane, G.W. Gabrielsen, R.J.G. Leakey, O.J. Lønne, M. Zajaczkowski, S. Falk-Petersen, M. Kendall, S.-Å. Wängberg, K. Bischof, A.Y. Voronkov, N.A. Kovaltchouk, J. Wiktor, M. Poltermann, G. di Prisco, C. Papucci, and S. Gerland. 2002. The marine ecosystem of Kongsfjorden, Svalbard. Polar Research 21: 167-208.
  106. Hop, H., K. Borgå, G.W. Gabrielsen, L. Kleivane, and J. U. Skaare. Food web magnification of persistent organic pollutants in poikilotherms and homeotherms from the Barents Sea food web. Environmental Science and Technology 36: 2589-2597.
  107. Engelsen, O., E.N. Hegseth, Hop, E. Hansen, and S. Falk-Petersen. 2002. Spatial variability of chlorophyll-a in the Marginal Ice Zone of the Barents Sea, with relations to sea ice and oceanographic conditions. Journal of Marine Systems 35: 79-97.
  108. Falk-Petersen, S., T.M. Dahl, C.L. Scott, , J.R. Sargent, B. Gulliksen, S. Kwasniewski, Hop, and R.-M. Millar. 2002. Lipid biomarkers and trophic linkages between the Arctic ctenophores and calanoid copepods in Svalbard waters. Marine Ecology Progress Series 227: 187-194.
  109. Gontar, V.I., Hop, and A. Yu. Voronkov. 2001. Diversity and distribution of Bryozoa in Kongsfjorden, Svalbard. Polish Polar Research 22: 187-204.
  110. Falk-Petersen, S., Hop, W.P. Budgell, E.N. Hegseth, R. Korsnes, T.B. Løyning, J.B. Ørbæk, T. Kawamura, and K. Shirasawa. 2000. Physical and ecological processes in the Marginal Ice Zone of the northern Barents Sea during the summer melt period. Journal of Marine Systems 27: 131-159.
  111. Poltermann, M., Hop, and S. Falk-Petersen. 2000. Life under Arctic sea ice – reproduction strategies of two sympagic (ice-associated) amphipod species, Gammarus wilkitzkii and Apherusa glacialis. Marine Biology 136: 913-920.
  112. Hop, H., M. Poltermann, O.J. Lønne, S. Falk-Petersen, R. Korsnes, and W.P Budgell. 2000. Ice-amphipod distribution relative to ice density and under-ice topography in the northern Barents Sea. Polar Biology 23: 357-367.
  113. Falk-Petersen, S., G. Pedersen, S. Kwasniewski, E.N. Hegseth, and Hop. 1999. Spatial distribution and life-cycle timing of zooplankton in the marginal ice zone of the Barents Sea during the summer melt season in 1995. Journal of Plankton Research 21: 1249-1264.
  114. Scott, C.L., S. Falk-Petersen, J.R. Sargent, Hop, O.J. Lønne, and M. Poltermann. 1999. Lipids and trophic interactions of ice fauna and pelagic zooplankton in the marginal ice zone of the Barents Sea. Polar Biology 21: 65-70.
  115. Falk-Petersen, S. J.R. Sargent, J. Henderson, E.N. Hegseth, Hop, and Y.B. Okolodkov. 1998. Lipids and fatty acids in ice algae and phytoplankton from the Marginal Ice Zone in the Barents Sea. Polar Biology 20: 41-47.
  116. Hop, H. and W.M. Tonn. 1998. Gastric evacuation rates and daily rations of Arctic cod (Boreogadus saida) at low temperatures. Polar Biology 19: 293-301.
  117. Hop, H., W.M. Tonn, and H.E. Welch. 1997. Bioenergetics of Arctic cod (Boreogadus saida) at low temperatures. Canadian Journal of Fisheries and Aquatic Sciences 54: 1772-1784.
  118. Hop, H., H.E. Welch, and R.E. Crawford. Population structure and feeding ecology of Arctic cod schools in the Canadian High Arctic. Pp. 68-80 In: J.B. Reynolds (ed.). Fish ecology in Arctic North America. American Fisheries Society Symposium 19, Bethesda, Maryland.
  119. Graham, M. and Hop. 1995. Aspects of reproduction and larval biology of Arctic cod (Boreogadus saida).  Arctic 48: 130-135.
  120. Hop, H. and M. Graham. Respiration of juvenile Arctic cod (Boreogadus saida): effects of acclimation, temperature, and food intake. Polar Biology 15: 359-367.
  121. Hop, H., M. Graham, and V.L. Trudeau. Spawning energetics of Arctic cod (Boreogadus saida) in relation to seasonal development of the ovary and plasma sex steroid levels. Canadian Journal of Fisheries and Aquatic Sciences 52: 541-550.
  122. Hop, H., J. Gjøsæter, and D.S. Danielssen. 1994. Dietary composition of sympatric juvenile cod, Gadus morhua, and juvenile whiting, Merlangius merlangus L., in a fjord of southern Norway. Aquaculture and Fisheries Management 25 (suppl.1): 49-64.
  123. Hop, H., D.S. Danielssen, and J. Gjøsæter. 1993. Winter feeding ecology of cod (Gadus morhua) in a fjord of southern Norway. Journal of Fish Biology 43: 1-18.
  124. Welch, H.E., R.E. Crawford, and Hop. 1993. Occurrence of Arctic cod (Boreogadus saida) schools and their vulnerability to predation in the Canadian high arctic. Arctic 46: 331-339.
  125. Hop, H., J. Gjøsæter, and D.S. Danielssen. 1992. Seasonal feeding ecology of cod (Gadus morhua) on the Norwegian Skagerrak coast. ICES Journal of Marine Science 49: 453-461.
  126. Welch, H.E., M.A. Bergmann, T.D. Siferd, K.A. Martin, M.F. Curtis, R.E. Crawford, R.J. Conover, and Hop. 1992. Energy flow through the marine ecosystem of the Lancaster Sound region, Arctic Canada. Arctic 45: 343-357.
  127. West, R.L., M.W. Smith, W.E. Barber, J.B. Reynolds, and Hop. 1992. Autumn migration and overwintering of Arctic grayling in coastal streams of the Arctic National Wildlife Refuge, Alaska. Transactions of the American Fisheries Society 121: 709-715.
  128. Hop, H. Fiskeørn Pandion h. haliaetus i Aust-Agder, Sør Norge, hekkebestand og reproduksjon 1980-88.  (Osprey Pandion h. haliaetus in Aust-Agder, southern Norway; breeding population and reproduction 1980-88.)  Fauna 42: 1-12. (In Norwegian with English summary).
  129. Hop, H. and A.J. Gharrett. 1989. Genetic relationships of Arctic grayling in the Koyukuk and Tanana rivers, Alaska. Transactions of the American Fisheries Society 118: 290-295.
  130. Armstrong, R.H., Hop, and J.H. Triplehorn. 1986. Indexed bibliography of the holarctic genus Thymallus (grayling) to 1985. Biological Papers of the University of Alaska 23: 19-110.

Popular science and outreach:

  • Plastics in the ocean. Chronicle in Nordlys 30 Dec., 2016
  • Kongsfjorden Ecosystem Workshop 2014 – new views after more than a decade of research. Ny-Ålesund Newsletter 34th Edition, July 2014.
  • Fishing for polar cod. Hop & von Quillfeldt. Chronicle in Nordlys 29 Jan., 2014.
  • Life below sea ice, in water masses and on the bottom around Svalbard. Tromstun School, 30 Jan., 2014
  • Contributed pictures (front/back cover) to book: Armstrong, R.H. and M.F. Wilson. 2014.
  • Natural connections in Alaska. Nature Alaska Images, Juneau, AK.
  • The ice edge-a sensitive site with much life. Interview, article in Barents Watch, 7 Jan., 2014
  • Deadly trio threatens the ocean. Interview with 2oC magazine, Nov., 2013.
  • Diving collection of organisms in Arctic sea ice. Interview on video, http://www.nrk.no/nordland, Oct., 2013.
  • The mackerel has arrived in Svalbard. Chronicle in Nordlys, 2 Oct., 2013.
  • When the power went in the freezer. Hop & Gerland. Chronicle in Nordlys, 28 Sept., 2013.
  • Arctic marine ecology and climate warming.  Interview with Radio France Internationale, 17 June, 2013
  • Move of die out. Consequences of ice-free ocean for life in the Arctic. Dagens Næringsliv, 24 Nov., 2013
  • What is Calanus?  Interview with NRK-P1, 17 August, 2012
  • Effects of climate change on Arctic marine ecosystems. Interview with Dokufilm, 17 Nov., 2010
  • ICE-research.  NRK Dagsrevyen, Nordnytt, 10 September, 2010
  • Arctic Marine Biology. Skjelnan skole, Tromsø, 28 May, 2010
  • Arctic Marine Biodiversity at Arctic Frontiers, Tromsø. “Verdt å vite”, NRK P2, 29 Jan., 2010
  • Followed climate change from the first row. Fakta fra Norklima 4, Oct. 2009
  • Kongsfjorden in Svalbard – passed the tipping point. Klima 6, 2009
  • Effects of climate warming in Svalbard fjords “Verdt å vite”, NRK P2, 15 Sept., 2009
  • Soon mackerel in the Arctic. Researchers fear loss of Arctic species. Interview, Aftenposten, 29 July, 2008
  • In Nature: On thin ice. Underwater filming of ice-associated organisms, NRK1, 25 Sept., 2007
  • Interview with VG about polar cod experiments in Kings Bay Marine Lab, Ny-Ålesund, 19 Sept., 2007
  • Interview article about climate change and Kongsfjorden in VEJA, Brazilian newspaper, 11 April, 2007
  • Arctic Society (Arktisk Forening) presentation on marine research under Arctic sea ice, 8 March, 2006
  • New Arctic marine laboratory in Ny-Ålesund, Svalbard. Nytt fra Nord 7, 2006
  • Arctic marine research with enhanced focus in Tromsø. Chronicle in Nordlys, 21 Sept., 2005
  • Article about the Arctic Marine Laboratory, Ny-Ålesund, interview in Morgenbladet, 27 May, 2005
  • Sea ice and climate warming; consequences for Arctic marine life, “Schrödingers katt” NRK1, 28 Oct., 2004
  • Marine biological research in Kongsfjorden: Kongsfjord Ecosystem Workshop II.
  • Ny-Ålesund Newsletter, June 2002
  • Pollutants in marine food webs in the Arctic, NRK Tromsø P2, 25 Sept., 2002
  • The rich forests and fauna of icy seas – new perspectives from diving in the Arctic.
  • Svalbardseminar – lecture series for the people of Longyearbyen, UNIS, 6 Feb., 2002
  • New information about life at the ice edge. Braatens Magazine 1, 2000
  • Interview and underwater film about sea ice ecology, “Schrödingers Katt” NRK1, 12 Oct., 2000
  • Polar bear eating ringed seal.  Cover photo for Natur & Miljø Bulletin 1-2, 2000
  • “Ishavets frodige skoger”, about diving project in Kongsfjorden. Svalbardposten 4, 1999
  • Interview about research in Kongsfjorden, Svalbard, “Verdt å vite”, NRK P2, 5 Sept., 2000
  • Interview about research cruise in the Marginal Ice Zone of the Barents Sea, NRK P1, 4 July, 1995
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