Here you can find a list of all ARCTOS members and PhD students. In the drop-down menu you can either select the member status or the institution the member belongs to.

Photo of Jasmine Nahrgang

Jasmine Nahrgang

Associate Professor

Research interests:
  • Impact of petroleum contaminants on Arctic marine organisms.
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UiT The Arctic University of NorwayArctic and Marine Biology Work Department of Arctic and Marine Biology Faculty of Biosciences, Fisheries and Economics UiT The Arctic University of Norway Tromsø 9037 Norway Work Phone: +47 77 64 58 96 Website: ORCID Website: ResearchGate Website: UiT researcher profile

Research interests:

  • Impact of petroleum contaminants on Arctic marine organisms.

Running national and international projects:

  • Participant – OCEANSPACT «OCEAN Sustainability Pathways for Achieving Conflict Transformation», Belmont Forum (€ 1.448.000), 2020-2022.
  • PI- PARENTox “Effects of parental exposure to petroleum on fertilization success and embryo development in Atlantic cod”, Fram Centre Flagship “MIKON” (kr 1.000.000,-) 2020 and 500.000,- from ARCEx.
  • Participant – NOR-LANKA Blue,  exchange programme with Sri Lanka. Norwegian Agency for International Cooperation and Quality Enhancement in Higher Education (DIKU) NORPART Programme, (kr 000.000,-), 2019-2023.
  • Participant – Arctic EcoSens “Environmental monitoring of Arctic coastal ecosystems: Sensitivity to petroleum pollution”, Norwegian Research Council, NORRUSS programme, (kr 10.000.000,-), 2018-2020.
  • Participant – Nansen Legacy, Research Focus “Human Impacts”, task “Effects of oil and contaminants on northern Barents Sea ecosystem health », (kr 000.000,-), 2018-2024.

Institutional responsibilities:

2019 –                   Coordinator for the Department´s Master programme in Biology

2019 –                   Member of the Board of Havbrukstasjon i Tromsø

2015 –                   Member of the Strategic Board for Education (programmestyret) of  the Department Arctic and Marine Biology, UiT.

Peer reviewed publications:

Annotations refer to:  *Main author (first, second or last) and † my students

  1. Meador JP, Nahrgang J* (2019) Characterizing Crude Oil Toxicity to Early-Life Stage Fish Based On a Complex Mixture: Are We Making Unsupported Assumptions? Environ Sci Technol 53:11080-11092.
  2. Nahrgang J*, Bender ML†, Meier S, Nechev J, Berge J, Frantzen M (2019) Growth and metabolism of adult polar cod (Boreogadus saida) in response to dietary crude oil. Environ. Saf. 180:53–62.
  3. Reinardy HC, Pedersen KB, Nahrgang J, Frantzen M (2019). Effects of mine tailings exposure on early life stages of Atlantic cod. Environ Toxicol Chem 38:1446–1454.
  4. Storhaug E†, Nahrgang J*, Pedersen KB, Brooks SJ, Petes LP, Bakhmet IN, Frantzen M (2019) Seasonal and spatial variations in biomarker baseline levels within Arctic populations of mussels (Mytilus spp.). Sci Tot Environ 656:921–936.
  5. Song Y, Nahrgang J*, Tollefsen K-E (2019) Transcriptomic analysis reveals dose-dependent modes of action of benzo(a)pyrene in polar cod (Boreogadus saida). Sci Tot Environ 653:176–189.
  6. Dahlke TF, Butzin M, Nahrgang J, Puvanendran V, Mortensen A, Pörtner HO, Storch D (2018) Northern cod species face spawning habitat losses if global warming exceeds 1.5°C. Science Advances 4 (11), eaas8821.
  7. Bender ML†, Frantzen M, Camus L, Le Floch S, Palerud J, Nahrgang J* (2018) Effects of acute exposure to dispersed oil and burned oil residue on long-term survival, growth, and reproductive development in polar cod (Boreogadus saida). Mar Environ Res 140:468–477.
  8. Fokina N, Storhaug E†, Bakhmet I, Maximovich N, Frantzen M, Nahrgang J* (2018) Seasonal changes in lipid class content in mussels Mytilus spp. from Rakkfjorden in the Norwegian Sea and Kandalaksha Bay of the White Sea. Polar Biol 41:2103–
  9. Vieweg I†, Bilbao E, Meador JP, Cancio I, Bender ML†, Cajaraville MP, Nahrgang J* (2018) Effects of dietary crude oil exposure on molecular and physiological parameters related to lipid homeostasis in polar cod (Boreogadus saida). Comp Biochem Physiol C 206-207:54–64.
  10. Vieweg I†, Benedetti M, Lanzoni I, Regoli F, Nahrgang J* (2017) Antioxidant defenses in polar cod (Boreogadus saida) and responsiveness towards dietary crude oil exposure. Mar Environ Res 130:48-59.
  11. Nahrgang J*, Dubourg P†, Frantzen M, Storch D, Dahlke F, Meador J (2016) Corrigendum to “Early life stages of an Arctic keystone species (Boreogadus saida) show high sensitivity to a water-soluble fraction of crude oil ». Pollut.
  12. Nahrgang J*, Dubourg P†, Frantzen M, Storch D, Dahlke F, Meador J (2016) Early life stages of an Arctic keystone species (Boreogadus saida) show high sensitivity to a water-soluble fraction of crude oil. Environ Pollut 218:605-614.
  13. Bender ML†, Frantzen M, Vieweg I†, Falk-Petersen IB, Johnsen HK, Rudolfsen G, Tollefsen KE, Dubourg P, Nahrgang J* (2016) Effects of chronic dietary petroleum exposure on reproductive development in polar cod (Boreogadus saida). Aquat Toxicol 180:196-208.
  14. Sagerup K, Nahrgang J*, Frantzen M, Larsen LH, Geraudie P (2016) Biological effects of marine diesel oil exposure in red king crab (Paralithodes camtschaticus) assessed through a water and foodborne exposure experiment. Mar Environ Res 119: 126-135.
  15. Mueter FJ, Nahrgang J*, Nelson RJ, Berge J (2016) The ecology of gadid fishes in the circumpolar Arctic with a special emphasis on the polar cod (Boreogadus saida). Polar Biol 39:961-967.
  16. Frantzen M, Regoli F, Ambrose WG, Nahrgang J, Geraudie P, Benedetti M, Locke W, Camus L (2016) Biological effects of mechanically and chemically dispersed oil on the Icelandic scallop (Chlamys islandica). Ecotoxicol Environ Saf 127:95-107.
  17. Nahrgang J*, Korshunova E†, Murzina SA, Delmas O, Nemova NN, Berge J (2016) Aspects of reproductive biology of wild caught polar cod (Boreogadus saida) from Svalbard waters. Polar Biol 39:1155-1164.
  18. Bakke M, Nahrgang J*, Ingebrigtsen K (2016) Comparative absorption and tissue distribution of 14C-benzo(a)pyrene and 14C-phenantrene in the polar cod (Boreogadus saida) following oral administration. Polar Biol 39:1165-1173.
  19. Berge J, Cottier F, Darnis G, Falk-Petersen S, Gabrielsen T, Johnsen G, Last K, Leu E, Lønne OJ, Moline M, Nahrgang J, Renaud PE, Seuthe L, Søreide J, Varpe Ø, Weslawski JM (2015) In the dark: paradigms of Arctic ecosystems during polar night challenged by new understanding. Progr Oceanogr 139:258-271.
  20. Camus L, Brooks S, Geraudie P, Hjorth M, Nahrgang J, Olsen GH, Smit MG (2015) Comparison of produced water toxicity to Arctic and temperate species. Ecotoxicol Environ Saf 113:248-58.
  21. Geraudie P, Nahrgang J*, Leray J, Minier C, Camus L (2014) Endocrine disrupting effects of produced water in polar cod (Boreogadus saida). J Toxicol Environ Health A 77:557-73.
  22. Nahrgang J*, Varpe Ø, Korshunova E†, Murzina S Hallanger IG, Vieweg I† (2014) Gender specific reproductive strategies of an Arctic key species (Boreogadus saida) and implications of climate change. PLOS One
  23. Berge J, Nahrgang J* (2013) The Atlantic spiny lumpsucker Eumicrotremus spinosus: life history traits and the seemingly unlikely interaction with the pelagic amphipod Themisto libellula. Polish Polar Res 34:279-287.
  24. Nahrgang J., Brooks S., Evenset A., Camus L., Jonsson M., Smith T., Lukina J., Frantzen M., Giarratano E., Renaud PE., 2013. Seasonal variation in biomarkers in blue mussel (Mytilus edulis), Icelandic scallop (Chlamys islandica) and Atlantic cod (Gadus morhua) – implications for environmental monitoring in the Barents Sea. Aquat. Toxicol. 127:21-35
  25. Frantzen M., Falk-Petersen I.B., Nahrgang J., Smith T., Olsen G.H., Hangstad T.A., Camus L., 2012. Toxicity of Goliat crude oil and pyrene to the embryos of beach spawning capelin (Mallotus villosus). Aquat. Toxicol. 108:42-52.
  26. Renaud P.E., Berge J., Varpe Ø., Lønne O.J., Nahrgang J., Ottesen C., Hallanger I., 2012. Climate change, competition, and consequences for the key Arctic species, Boreogadus saida. Polar Biol. 35:401-412.
  27. Camus L., Nahrgang J., Olsen G.H., 2010. Risk assessment and biomonitoring tools in the Arctic. Proceedings of 33rd AMOP Technical Seminar on Environmental Contamination and Response, Halifax, Canada. Volume 2, 1253-1266
  28. Nahrgang J., Camus L., Broms F., Christiansen J.S., Hop H., 2010. Seasonal baseline levels of physiological and biochemical parameters in polar cod (Boreogadus saida): implications for environmental monitoring. Mar. Pollut. Bull. 60:1336-1345
  29. Nahrgang J., Jönsson M., Camus L., 2010. EROD activity in liver and gills of polar cod (Boreogadus saida) exposed to waterborne and dietary crude oil. Mar. Environ. Res. 70:120-123
  30. Christiansen J.S., Karamushko L.I., Nahrgang J., 2010. Sub-lethal levels of waterborne petroleum may depress routine metabolism in polar cod Boreogadus saida (Lepechin, 1774). Polar Biol. 33:1049-1055
  31. Nahrgang J., Camus L., Carls M.G., Gonzalez P., Jönsson M., Taban I.C., Bechmann R.K., Christiansen J.S., Hop H. 2010. Biomarker responses in polar cod (Boreogadus saida) exposed to the water soluble fraction of crude oil. Aquat. Toxicol. 97:234-242
  32. Nahrgang J., Camus L., Gonzalez P., Jönsson M., Christiansen J.S., Hop H. 2010. Biomarker responses in polar cod (Boreogadus saida) exposed to dietary crude oil. Aquat. Toxicol. 96:77-83
  33. Nahrgang J., Camus L., Gonzalez P., Goksøyr A., Christiansen J.S., Hop H., 2009. PAH biomarker responses in polar cod (Boreogadus saida) exposed to benzo(a)pyrene. Aquat. Toxicol. 94:309-319
Photo of Fredrika Norrbin

Fredrika Norrbin

Associate Professor

Research interests:
  • Zooplankton vertical distribution
  • Life cycles
  • Overwintering strategies and phenology
  • Biogeography.
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UiT The Arctic University of NorwayDepartment of Arctic and Marine Biology Work Department of Arctic and Marine Biology Faculty of Biosciences, Fisheries and Economics UiT The Arctic University of Norway Tromsø 9037 Norway Work Phone: +47 77 64 47 47

Research interests:

  • Zooplankton vertical distribution
  • Life cycles
  • Overwintering strategies and phenology
  • Biogeography.

Layers and aggregations of zooplankton in relation to environmental parameters. To what degree is vertical distribution affected by biological drivers; e.g. competition from transient species and predators, compared to physical parameters. Phenology and timing of life history event; overwintering strategies, timing of diapause (cont. of PhD). Biogeography of small copepods, especially Pseudocalanus sibling species. Plankton investigations in North Norwegian fjords, Arctic and Atlantic regions, where identifying distributions with the VPR is a vital part.

Completed national and international projects:

  • Pelagic ecosystems in ice-covered and ice-free fjords under climate change (Fram Centre, Coast & Sea Flagship, 2013-2015)
  • Analysis of Video Plankton Recorder Data, Iceland-Norway workshops (2014)
  • Pseudocalanus: ecology and taxonomy of two sibling copepod species in Svalbard and Northern Norwegian Waters (2009-2013)
  • The significance of thin layers in aquatic plankton communities (NRC, 2005.2009)
  • Autonomous VPR system. Vertically profiling video plankton recording system with simultaneous collection of CTD data and fluorescence (NRC, 2002-2004)

Relevant publications:

  1. Aarbakke, Ole Nicolai S; Bucklin, Ann; Halsband, Claudia; Norrbin, Fredrika. Comparative phylogeography and demographic history of five sibling species of Pseudocalanus (Copepoda: Calanoida) in the North Atlantic Ocean. Journal of Experimental Marine Biology and Ecology 2014; Volum 461. ISSN 0022-0981.s 479 – 488.s doi: 10.1016/j.jembe.2014.10.006.
  2. Basedow, Sünnje; Tande, Kurt; Norrbin, Fredrika; Kristiansen, Stian Andre. Capturing quantitative zooplankton information in the sea: Performance test of laser optical plankton counter and video plankton recorder in a Calanus finmarchicus dominated summer situation. Progress in Oceanography 2013; Volum 108. ISSN 0079-6611.s 72 – 80.s doi: 10.1016/j.pocean.2012.10.005.
  3. Aarbakke, Ole Nicolai S; Bucklin, Ann; Halsband, Claudia; Norrbin, Fredrika. Discovery of Pseudocalanus moultoni (Frost, 1989) in Northeast Atlantic waters based on mitochondrial COI sequence variation. Journal of Plankton Research 2011; Volum 33 (10). ISSN 0142-7873.s 1487 – 1495.s doi: 10.1093/plankt/fbr057.
  4. Jacobsen, Helge Petter; Norrbin, Fredrika. Fine-scale layer of hydromedusae is revealed by video plankton recorder (VPR) in a semi-enclosed bay in northern Norway. Marine Ecology Progress Series 2009; Volum 380. ISSN 0171-8630.s 129 – 135.s doi: 10.3354/meps07954.
  5. Norrbin, Fredrika; Eilertsen, Hans Christian; Degerlund, Maria Ingeborg. Vertical distribution of primary producers and zooplankton grazers during different phases of the Arctic spring bloom. Deep-sea research. Part II, Topical studies in oceanography 2009; Volum 56 (21-22). ISSN 0967-0645.s 1945 – 1958.s doi: 10.1016/j.dsr2.2008.11.006.
  6. Wold, A; Norrbin, Fredrika. Vertical migration as a response to UVR stress in Calanus finmarchicus females and nauplii. Polar Research 2004; Volum 23 (1). ISSN 0800-0395.s 27 – 34.
  7. Norrbin, Fredrika. Ultra-structural changes in overwintering females of Acartia longiremis Lilljeborg. Marine Biology 2001; Volum 139. ISSN 0025-3162.s 697 – 704.
  8. Norrbin, Fredrika; Davis, Cabell S.; Gallager, Scott. Differences in fine-scale structure and composition of zooplankton between mixed and stratified regions of George’s Bank. Deep-sea research. Part II, Topical studies in oceanography 1996; Volum 43. ISSN 0967-0645.s 1905 – 1924.
  9. Norrbin, Fredrika. Timing of diapause in relation to the onset of winter in the high-latitude marine copepods Pseudocalanus acuspes and Acartia longiremis. Marine Ecology Progress Series 1996; Volum 142. ISSN 0171-8630.s 99 – 109.
  10. Hansen, Benni W; Verity, Peter; Falkenhaug, Tone; Tande, Kurt Steinar; Norrbin, Fredrika. On the trophic fate of Phaeocystis pouchetii (Harriot). V. Trophic relationships between Phaeocystis and zooplankton: an assessment of methods and size dependence. Journal of Plankton Research 1994; Volum 16. ISSN 0142-7873.s 487 – 511.
  11. Norrbin, Fredrika. Seasonal patterns in gonad maturation, sex ratio and size in some small, high latitude copepods: implications for overwintering tactics. Journal of Plankton Research 1994; Volum 16 (2). ISSN 0142-7873.s 115 – 131.
  12. Norrbin, Fredrika. Skewed sex ratios in overwintering copepodites of Pseudocalanus in Norwegian fjords – a result of sex-specific selection?. Bulletin of marine science 1993; Volum 53. ISSN 0007-4977.s 204 – 215.
  13. Noji, Thomas; Estep, Kenneth; McIntyre, Ferren; Norrbin, Fredrika. Image analysis of faecal material grazed upon by three species of copepods: evidence for coprorhexy, coprophagy and coprochaly. Journal of the Marine Biological Association of the United Kingdom 1991; Volum 71. ISSN 0025-3154.s 465 – 480.
  14. Norrbin, Fredrika. Gonad maturation as an indication of seasonal cycles for several species of small copepods in the Barents Sea. Polar Research 1991; Volum 10. ISSN 0800-0395.s 421 – 432.
  15. Norrbin, Fredrika; Olsen, Rolf Erik; Tande, Kurt Steinar. Seasonal variation in lipid class and fatty acid composition of two small copepods in Balsfjorden, Northern Norway. Marine Biology 1990; Volum 105. ISSN 0025-3162.s 205 – 211.
  16. Norrbin, Fredrika; Båmstedt, Ulf. Energy contents in benthic and planktonic invertebrates of Kosterfjorden, Sweden. A comparison of energetic strategies in marine organism groups. Ophelia : International Journal of Marine Biology 1984; Volum 23 (1). ISSN 0078-5326.s 47 – 64.

Popular science and outreach:

  • Norrbin, F. (2014) A Focus on Plankton. Fram Forum 2014
  • Norrbin, F. (2013) Blog entry, PAP-cruise (Porcupine Abyssal Plains) . Imaging twilight critters.
  • Forskningsdagene 2008, organized activities on R/V Johan Ruud, two days
Photo of Robynne Calypso Nowicki

Robynne Calypso Nowicki

PhD candidate

PhD project title: Seasonal variation of energy and pollutants in the Barents Sea pelagic food web.…Read More
UNIS – The University Centre in SvalbardArctic Biology Work UNIS – The University Centre in Svalbard Post box 156 Longyearbyen 9171 Svalbard Website: ResearchGate

PhD project title:

Seasonal variation of energy and pollutants in the Barents Sea pelagic food web.

PhD project description:

This study aims to quantify the seasonal changes in species specific energy content, using bomb calorimetry, within and across trophic levels (macrozooplankton and fish). We aim to assess potential “critical windows”, i.e. when organisms are in a sub-optimal state and therefore more susceptible to stress, such persistent organic pollutants (POPs). We will also evaluate how seasonal energy content and pollutant levels of prey will impact the top-predators in the system, particularly seabirds. In addition, tissue specific samples will be taken for POP concentration, to study seasonal remobilization in key marine organisms.

Start date: June 2019
Planned submission date: June 2022


Øystein Varpe (UiB, Bergen, Norway)
Katrine Borgå (UiO, Oslo, Norway)
Geir W. Gabrielsen (NPI, Tromsø, Norway)
Janne Søreide (UNIS, Longyearbyen, Svalbard)

Scientific publications:

  1. David Palma, Alix Varnajot, Kari Dalen, Ilker K. Basaran, Charles Brunette, Marta Bystrowska, Anastasia D. Korablina, Robynne C. Nowicki & Thomas A. Ronge (2019) Cruising the marginal ice zone: climate change and Arctic tourism, Polar Geography, 42:4, 215-235, DOI: 10.1080/1088937X.2019.1648585


  • Head of Education and Outreach, UK Polar Network
  • Longyearbyen event organizer, Pint of Science Norway