Tore Haug » Members

Here you can find a list of all ARCTOS members and PhD students. In the drop-down menu you can either select the member status or the institution the member belongs to.

Tore Haug

Researcher and research group leader (IMR) Institute of Marine ResearchMarine mammals group
Work address Institute of Marine Research, Tromsø Post box 6404 Tromsø 9294 Norway Work Phone: +47 95 28 42 96
Photo of Tore Haug

Research interests:

  • Questions related to management and conservation of whale and seal stocks in Norwegian and adjacent waters
  • Distribution abundance productivity harvest potential and their role in the ecosystem
  • Responsible for IMRs’ advice to Norwegian authorities on whale and seal stocks in Norwegian waters.

Running national and international projects:

IMR (Havforskningsinstituttet):
2005-: Forvaltning av ishavssel
2005-: Sjøpattedyrenes økologiske betydning
2005-: DNA-registeret vågekval
2010-: Fra plankton til sel (FRAM-prosjekt)

Completed national and international projects:

Research Council of Norway (Norges Forskningsråd):

  • Responsible for the following projects:
  • 1990-1994: ‘Ernæring og kondisjon hos grønlandssel i Barentshavet (Feeding and body condition in Barents Sea harp seals)’ (4001-2800.083)
  • 1990-1994: ‘Ernæringsøkologi hos nordøst-atlantisk vågekval (Feeding ecolpogy of Northeast Atlantic minke whales’ (104499/110), including the research whaling under scientific permit.
  • 1995-1997: ‘Grønlandsselens rolle i Barentshavets flerbestandssystem (The role of harp seals in the multspecies interactions of the Barents Sea)’ (108147/110).
  • 1995-1997: ‘Metodeutvikling for overvåkning av vågekvalens matkonsum i Barentshavet (Development of methods to monitor the consumption by minke whales in the Bartnets Sea)’ (108146/110).
  • 1997-2000:’Prey preferences in minke whales’ (113485/122).
  • 1998-1999:’Prey selectivity and responses to changes in prey abundance of minke whales’ (121385/122).
  • 2000-2002:’Ecological role of harp and hooded seals in the Nordic Seas’ (133646/122)
  • 2001-2002:’Prediction models of the impact of minke whales and harp seals on fish stocks, postdoc Ulf Ove Lindstrøm (140554/120)’
  • 2002:’Improving assessment and ecology research on harp and hooded seals, Guest scientist scholarship (146573/120)’
  • 2014-2018: SI Arctic (nestleder)

Curriculum vitae: PDF

Selected publications (last 10 years):

  1. Stenson, G.B., Haug, T. & Hammill, M.O. 2020. Harp Seals: Monitors of Change in Differing Ecosystems. Frontiers in. Marine Science 7:569258. Doi:10.3389/fmars.2020.569258
  2. Gunnlaugsson, T., Vikingsson, G.A., Halldorsson, S.D., Elvarsson, B.Þ., Haug, T. & Lydersen, C. 2020. Body mass, muscle, blubber and visceral fat content and their seasonal, spatial and temporal variability in North Atlantic common minke whales. Journal of Cetacean Research and Management 21: 59–70.
  3. Nilssen, K.T., Lindstrøm, U., Westgaard, J.I., Lindblom, L., Blencke, T.-R. & Haug, T. 2019. Diet and prey consumption of grey seals (Halichoerus grypus) in Norway. Marine Biology Research 15: 137-149. Doi: 10.1080/17451000.2019.1605182.
  4. Moore, S.E., Haug, T., Vikingsson, G.A.&d Stenson, G.B. 2019.Baleen whale ecology in arctic and subarctic seas in an era of rapid habitat alteration. Progress in Oceanography 176: 102118. 15 pp.
  5. Nakamura, G., Ryeng, K.A., Kadowaki, I., Hayashi, R., Nagatsuka, S., Hirose, A., Fujise, Y. and Haug, T. 2018. Comparisons of shapes of the white flipper patch between two subspecies of common minke whales (Balaenoptera acutorostrata). Cetacean Population Studies 1: 15-24.
  6. Vacquié-Garcia, J., Lydersen, C., Biuw, M., Haug, T., Fedak, M.A. Kovacs, K.M. 2017. Hooded seal Cistophora cristata foraging areas in the Northeast Atlantic Ocean – investigated using three complementary methods. Plos One 12(12): e0187889.
  7. Enoksen, S., Haug, T., Lindstrøm, U. & Nilssen, K.T. 2017. Recent summer diet of hooded Cystophora cristata and harp Pagophilus groenlandicus seals in the drift ice of the Greenland Sea. Polar Biology 40: 931-937. Doi 10.1007/s00300-016-2002-2.
  8. Haug, T., Falk-Petersen, S., Greenacre, M., Hop, H., Lindstrøm, U., Meier, S., Nilssen, K.T. and Wold, A. 2017. Trophic level and fatty acids in harp seals compared with common minke whales in the Barents Sea. Marine Biology Research 13: 919-932.
  9. Haug, T., Bogstad, B., Chierici, M., Gjøsæter, H., Hallfredsson, E.H., Høines, Å.S., Hoel, A.H., Ingvaldsen, R.B., Jørgensen, L.L., Knutsen, T., Loeng, H., Naustvoll, L.-J., Røttingen, I. & Sunnanå, K. 2017. Future harvest of living resources in the Arctic Ocean of the Nordic and Barents Seas: A review of possibilities and constraints. Fisheries Research 188: 38-57.
  10. Solvang, H.K., Yanagihara, H., Øien, N. & Haug, T. 2017. Temporal and geographical variation in body condition of common minke whales (Balaenoptera acutorostrata acutorostrata) in the Northeast Atlantic. Polar Biology 40: 667-683. Doi 10.1007/s00300-016-1992-0.
  11. Malde, K., Seliussen, B.B., Quintela, M., Dahle, G., Besnier, F., Skaug, H.J., Øien, N., Solvang, H.K., Haug, T., Skern-Mauritzen, Kanda, N., Pastene, L.A., Jonassen, I., & Glover, K.A. 2017. Whole genome resequencing reveals diagnostic markers for investigating global migration and hybridization between minke whale species. BMC Genomics 18: 76. Doi 10.1186/s12864-016-3416-5.
  12. Yamamura, M., Yangihara, H., Solvang, H.K., Øien, N. & Haug, T. 2016. Canonical correlation analysis for geographical and chronological responses. Procedia Computer Science 96: 1351-1360. Doi:10.1016/j.procs.2016.08.180.
  13. Meier, S. Falk-Petersen, S., Gade-Sørensen, L.A., Greenacre, M., Haug, T. & Lindstrøm, U. 2016. Fatty acids in common minke whale (Balaenoptera acutorostrata) blubber reflect the feeding area and food selection, but also high endogenous metabolism. Mar. Biol. Res. 12: 221-238.
  14. Bogstad, B., Gjøsæter, H., Haug, T. & Lindstrøm, U. 2015. A review of the battle for food in the Barents Sea: cod vs. marine mammals. Frontiers Ecol. Evol. 3, 29. doi:10.3389/fevo.2015.0029.
  15. Quintela, M., Skaug, H.J., Øien, N., Haug, T., Seliussen, B., Solvang, H.K., Pampoulie, C., Kanda, N., Pastene, L.A. & Glover, K. 2014. Investigating population genetic structure in a highly mobile marine organism: the minke whales Balaenoptera acutorostrata acutorostrata in the north east Atlantic. PLoS ONE 9(9): e108640. doi: 10.1371/journal.pone.0108640.
  16. Øigård, T.A., Haug, T. & Nilssen, K.T. 2014. From pup production to quotas: Current status of harp seals in the Greenland Sea. ICES J. Mar. Sci. 71: 537-545. doi:10.1093/icesjms/fst155
  17. Øigård, T.A., Haug, T. & Nilssen, K.T. 2014. Current status of hooded seals in the Greenland Sea. Victims of climate change and predation? Biol. Conserv. 172: 29-36.
  18. Øigård, T.A., Lindstrøm, U., Haug, T., Nilssen, K.T. & Smout, S. 2013. Functional relationship between harp seal body condition and available prey in the Barents Sea. Mar. Ecol. Prog. Ser. 484: 287-301. doi 10.3354/meps10272
  19. Nymo, I.H., Tryland, M., Frie, A.K., Haug, T., Foster, G., Rødven, R. & Godfroid, J. 2013. Age-dependent prevalence of anti-Brucella antibodies in hooded seals Cystophora cristata. Dis. Aquat. Org. 106: 187-196. Doi: 10.3354/dao02659
  20. Lindstrøm, U., Nilssen, K.T., Pettersen, L.M.S. & Haug, T. 2013. Harp seal foraging behaviour during summer around Svalbard in the northern Barents Sea: diet composition and the selection of prey. Polar Biol. 36: 305-320. doi 10.1007/s00300-012-1260-x
  21. Iversen, M., Aars, J., Haug, T., Alsos, I.G., Lydersen, C., Bachmann, L. & Kovacs, K.M. 2013. The diet of polar bears (Ursus maritimus) from Svalbard, Norway, inferred from scat analysis. Polar Biol. 36: 561-571. doi 10.1007/s00300-012-1284-2
  22. Glover, K.A., Kanda, N., Haug, T., Pastene, L.A., Øien, N., Seliussen, B.B., Sørvik, A.G.E. & Skaug, H.J. 2013. Hybrids between common and Antarctic minke whales are fertile and can back-cross. BMC Genetics 14: 25 (11 pp.). doi: 10.1186/1471-2156-14-25.
  23. Leclerc, L.-M.E., Lydersen, C., Haug, T., Bachmann, L., Fisk, A.T. & Kovacs, K.M.. 2012. A missing piece in the Arctic food web puzzle? Stomach contents of Greenland sharks sampled in Svalbard, Norway. Polar Biol. 35: 1197-1208. Doi 10.1007/s00300-012-1166-7.
  24. Torrissen, O., Glover, K.A., Haug, T., Misund, O.A., Skaug, H.J. & Kaiser, M. 2012. Good ethics or political and cultural censoring in science? ICES J. Mar. Sci. 69: 493-497.
  25. Frie, A.K., Haug, T. & Stenson, G. 2012. Long term trends in reproductive and demographic parameters of female Northwest Atlantic hooded seals (Cystophora cristata): Population responses to ecosystem change? Can. J. Zool. 90: 376-392.
  26. Glover, K.A., Haug, T., Øien, N., Walløe, L., Lindblom, L., Seliussen, B.B. & Skaug, H.J. 2011. The Norwegian minke whale DNA register: a data base monitoring commercial harvest and ttrade of whale products. Fish Fisher. 13: 313-332. doi: 10.1111/j.1467-2979.2011.00447.x
  27. Leclerc, L.-M., Lydersen, C., Haug, T., Glover, K.A., Fisk, A.T. & Kovacs, K.M. 2011. Greenland sharks (Somniosus microcephalus) scavenge offal from minke (Balaenoptera acutorostrata) whaling operations in Svalbard (Norway). Polar Res. 30, 7342, 4 pp. DOI: 10.342/polar.v30i0.7342.
  28. Labansen, A.L., Lydersen, C., Levermann, N., Haug, T. & Kovacs, K.M. 2011. Diet of ringed seals (Pusa hispida) from northeast Greenland. Polar Biol. 34: 227-234.
  29. Grahl-Nielsen, O., Haug, T., Lindstrøm, U. & Nilssen, K.T.. 2011. Fatty acids in harp seal blubber do not necessarily reflect their diet. Mar. Ecol. Prog. Ser. 426: 263-273.
  30. Johansen, C.E., Lydersen, C., Aspholm, P.E., Haug, T. & Kovacs, K.M. 2010. Helminth parasites in ringed seals (Pusa hispida) from Svalbard, Norway, with special emphasis on nematodes: variation with age, sex and location of host. J. Parasitol. 96: 946-953.
  31. Skoglund, E.G., Lydersen, C., Grahl-Nielsen, O., Haug, T. & Kovacs, K.M. 2010. Fatty acid composition of the blubber and dermis of adult male Atlantic walruses (Odobaenus rosmarus rosmarus) in Svalbard, and their potential prey. Mar. Biol. Res. 6: 239-250.
  32. Øigård, T.A., Haug, T., Nilssen, K.T. and Salberg, A.B. 2010. Estimation of pup production of harp and hooded seals in the Greenland Sea in 2007: Reducing uncertainty using generalized additive models. J. Northw. Atl. Fish. Sci. 42: 103-123.
  33. Nilssen, K.T., Skavberg, N.E., Poltermann, M., Haug, T., Härkönen, T. & Henriksen, G. 2010. Status of harbour seals (Phoca vitulina) in mainland Norway. NAMMCO Sci. Publ. 8:61-69.
  34. Mogren, H.-G., Lindstrøm, U., Nilssen, K.T. & Haug, T. 2010. Haulout behaviour of harbour seals during breeding and moult in Vesterålen, Norway. NAMMCO Sci. Publ. 8: 267-276.
  35. Glover, K., Kanda, N., Haug, T., Pastene, L.A., Øien, N., Goto, M., Seliussen, B.B. & Skaug, H.J. 2010. Migration of Antarctic minke whales to the Arctic. PLoS ONE: 5 (12) e15197: 1-6.

Selected other publications:

  1. Haug, T., Bjørge, A., Øien, N. & Ziryanov, S.V. 2011. 7. Marine mammals; 7.1 Marine mammals of the Barents Sea. Pp 395-430 in: Jakobsen, T. & Ozhigin, V.K. (eds). The Barents Sea; Ecosystem, resources, management; Half a century of Russian-Norwegian cooperation. Tapir Academic Press. 825 pp.
  2. Haug, T., Skern-Mauritzen, M. & Lindstrøm, U. 2011. 8. Trophic relationships; 8.5 Predation by marine mammals. Pp 485-494 in: Jakobsen, T. & Ozhigin, V.K. (eds). The Barents Sea; Ecosystem, resources, management; Half a century of Russian-Norwegian cooperation. Tapir Academic Press. 825 pp.
  3. Haug, T., Øien, N. & Golyak, I. 2011. 10. Research surveys; 10.5 Marine mammal surveys. Pp 595-603 in: Jakobsen, T. & Ozhigin, V.K. (eds). The Barents Sea; Ecosystem, resources, management; Half a century of Russian-Norwegian cooperation. Tapir Academic Press. 825 pp.

Popular science and outreach (last five years):

  1. Haug, T., Øien, N., Skaug, H.J. & Glover, K. 2015. DNA-register for vågehval: en skattekiste for studier av hvalbiologi. Pp. 108-109 i Bakketeig, I.E., Gjøsæter, H., Hauge, M., Sunnset, B.H. & Toft, K.Ø. (Eds.) Havforskningsrapporten 2015, Fisken og havet, Særnummer 1-2015. Havforskningsinstituttet, Bergen.
  2. Haug, T. & Øigård, T.A. 2015. Sel – Grønlandssel & Klappmyss. Pp. 186-187 i Bakketeig, I.E., Gjøsæter, H., Hauge, M., Sunnset, B.H. & Toft, K.Ø. (Eds.) Havforskningsrapporten 2015, Fisken og havet, Særnummer 1-2015. Havforskningsinstituttet, Bergen.
  3. Haug, T. & Øigård, T.A. 2016. Sel – Grønlandssel & Klappmyss. Pp. 174-175 i Bakketeig, I.E., Hauge, M., Kvamme, C., Sunnset, B.H. & Toft, K.Ø. (Eds.) Havforskningsrapporten 2016, Fisken og havet, Særnummer 1-2016. Havforskningsinstituttet, Bergen.
  4. Haug, T. 2017. Sel – Grønlandssel & Klappmyss. Pp. 66-67 i Bakketeig, I.E., Hauge, M. & Kvamme, C. (Eds.) Havforskningsrapporten 2017, Fisken og havet, Særnummer 1-2017. Havforskningsinstituttet, Bergen.
  5. Haug, T. 2017. Klappmyss. Store Norske Leksikon, Artikkel, 10.november 2017.
  6. Haug, T. 2017. Grønlandssel. Store Norske Leksikon, Artikkel, 19.desember 2017.
  7. Haug, T., Bogstad, B., Ingvaldsen, R., Knutsen, T. & Naustvoll, L.-J. 2017. Et varmere Arktis skaper trøbbel for de lokale artene., kronikk, 05.07.2017.
  8. Haug, T. 2018. Sel – Grønlandssel & Klappmyss. Pp. 38-39 i Huse, G. & Bakketeig, I.E. (Eds.) Ressursoversikten 2018, Fisken og havet, 6-2018. Havforskningsinstituttet, Bergen.
  9. Haug, T. 2018. Vågehval. Store Norske Leksikon, Artikkel, 11.juni 2018.
  10. Haug, T. 2018. Finnhval. Store Norske Leksikon, Artikkel, 29.juni 2018.
  11. Haug, T. 2018. Blåhval. Store Norske Leksikon, Artikkel, 30.juli 2018.
  12. Haug, T. 2018. Bardehvaler. Store Norske Leksikon, Artikkel, 30.juli 2018.
  13. Haug, T. 2018. Knølhval. Store Norske Leksikon, Artikkel, 28.september 2018.
  14. Haug, T. 2019. Spermhval. Store Norske Leksikon, Artikkel, 9.april 2019.
  15. Haug, T. 2019. Seihval. Store Norske Leksikon, Artikkel, 5.juli 2019.
  16. Haug, T. 2019. Grønlandshval. Store Norske Leksikon, Artikkel, 5.juli 2019.
  17. Haug, T. 2019. Havert. Store Norske Leksikon, Artikkel, 10.juli 2019.
    Haug, T. 2019. Retthvaler. Store Norske Leksikon, Artikkel, 28.november 2019.
  18. Haug, T. 2019. Nordkaper. Store Norske Leksikon, Artikkel, 28.november 2019.
  19. Haug, T. 2020. Brydehval. Store Norske Leksikon (, Artikkel, 20.mars 2020.
  20. Haug, T. 2020. Steinkobbe. Store Norske Leksikon (, Artikkel. 24.mars 2020.
  21. Haug, T. 2020. Grønlandssel. Store Norske Leksikon (, Artikkel. 25.mars 2020.
  22. Haug, T. 2020. Blueback. Store Norske Leksikon (, Artikkel. 25.mars 2020.
  23. Haug, T. 2020. Spekkhogger. Store Norske Leksikon (, Artikkel. 8.april 2020.
  24. Haug, T. 2020. Nise. Store Norske Leksikon (, Artikkel. 9.september 2020.
  25. Haug, T. 2020. Spermhvaler. Store Norske Leksikon (, Artikkel. 21.september 2020.
  26. Haug, T. 2020. Finnhvaler. Store Norske Leksikon (, Artikkel. 21.september 2020.
  27. Haug, T. 2020. Nebbhvaler. Store Norske Leksikon (, Artikkel. 21.september 2020.
  28. Haug, T. 2020. Nebbhval. Store Norske Leksikon (, Artikkel. 9.oktober 2020.
  29. Haug, T. and Kildebo, N. 2020. Niser. Store Norske Leksikon (, Artikkel. 8.april 2020.