Here you can find a list of all ARCTOS members and PhD students. In the drop-down menu you can either select the member status or the institution the member belongs to.

Photo of Anette Wold

Anette Wold

Chief engineer

Research interests:
  • Plankton ecology with a main emphasis on phytoplankton and zooplankton
  • The impact of changing sea ice conditions and associated effects on phytoplankton and ice algal communities
  • Ecosystem structure and energy flow of Arctic marine food webs including fatty acid throphic markers and stable isotope studies.
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Norwegian Polar InstituteResearch department: ecotoxicology Work address Norwegian Polar Institute Fram Centre Tromsø 9296 Norway Work Phone: +47 77 75 05 46

Research interests:

  • Plankton ecology with a main emphasis on phytoplankton and zooplankton
  • The impact of changing sea ice conditions and associated effects on phytoplankton and ice algal communities
  • Ecosystem structure and energy flow of Arctic marine food webs including fatty acid throphic markers and stable isotope studies.

Running national and international projects:

  • The Nansen Legacy (2018-2022)
  • Arctic Ecosystem Monoitoring: Pelagic study in Kongsfjorden (2010->)
  • MOSJ: Pelagic study in Kongsfjorden (2009->)

Completed national and international projects:

  • N-ICE2015: Ecosystem dynamics in a high-Arctic pack ice environment (5 Jan 2015–30 Jun 2015)
  • CLEOPATRA II: Climate effects on planktonic food quality and trophic transfer in the Arctic marginal ice zone (1 Jul 2012–30 Jul 2015)

Relevant scientific publications:

  1. Hop H, Wold A, Vihtakari M, Daase M, Kwasniewski S, Gluchowska M, Lischka S, Buchholz F, Falk-Petersen S (accepted) Zooplankton in Kongsfjorden (1996-2016) in relation to climate change. Advances in Polar Ecology
  2. Darnis G, Wold A, Falk-Petersen S, Graeve M, Fortier L (2018) Could offspring predation offset the successful reproduction of the arctic copepod Calanus hyperboreus under reduced sea-ice cover conditions? Progress in Oceanography 170: 107:118.
  3. Fernández-Méndez M, Olsen LM, Kauko HM, Meyer A, Rösel A, Merkouriadi I, Mundy CJ, Ehn JK, Johansson MA, Wagner PM, Ervik Å, Sorrell BK, Duarte PM, Wold A, Hop H, Assmy P (2018) Algal hot spots in a changing Arctic Ocean: Sea-ice ridges and the snow-ice interface. Frontiers in Marine Science
  4. Brown TA, Assmy P, Hop H, Wold A, Belt S (2017) Transfer of ice algae carbon to ice associated amphipods in the high-Arctic pack ice environment. J. Plankton Res. 00(00): 1–11. doi:10.1093/plankt/fbx030
  5. Haug T, Falk-Petersen S, Greenacre M, Hop H, Lindstrøm U, Meier S, Nilssen KT, Wold A (2017). Trophic level and fatty acids in harp seals compares with common minke whales in the Barents Sea. Marin. Biol. Res.
  6. Darnis G, Hobbs L, Geoffroy M, Grenvald JC, Renaud PE, Berge J, Cottier F, Kristiansen S, Daase M, Søreide J, Wold A, Morata N, Gabrielsen T (2017). From polar night to midnight sun: Diel vertical migration, metabolism and biogeochemical role of zooplankton in a high Arctic fjord (Kongsfjorden, Svalbard). Limnol. Oceanogr., doi: 10.1002/lno.10519.
  7. Assmy P, Fernández-Méndez M, Duarte P, Meyer A, Randelhoff A, Mundy CJ, Olsen LM, Kauko HM, Bailey A, Chierici M, Cohen L, Doulgeris AP, Ehn JK, Fransson A, Gerland S, Hop H, Hudson SR, Hughes N, Itkin P, Johnsen G, King JA, Koch BP, Koenig Z, Kwasniewski S, Laney SR, Nicolaus M, Pavlov AK, Polashenski CM, Provost C, Rösel A, Sandbu M, Spreen G, Smedsrud LH, Sundfjord A, Taskjelle T, Tatarek A, Wiktor J, Wagner PM, Wold A, Steen H, Granskog MA (2017). Leads in Arctic pack ice enable early phytoplankton blooms below snow-covered sea ice. Scientific Reports 7:40850 DOI:10.1038/srep40850
  8. Fransson A, Chierici M, Hop H, Findlay HS, Kristiansen S, Wold A (2016) Late winter-to-summer change in ocean acidification state in Kongsfjorden, with implications for calcifying organisms. Polar Biology 39:1841:1857. DOI 10.1007/s00300-016-1955-5
  9. Dalpadado P, Hop H, Rønning J, Pavlov V, Sperfeld E, Buchholz F, Rey A, Wold A (2015) Distribution and abundance of euphausiids and pelagic amphipods in Kongsfjorden, Isfjorden and Rijpfjorden (Svalbard) and changes in their relative importance as key prey in a warming marine ecosystem. Polar Biol. DOI 10.1007/s00300-015-1874-x
  10. Mayzaud, P., Falk-Petersen, S., Noyon, M., Wold, A., Boutoute, M. 2015. Lipid composition of three co-existing Calanus species in the Arctic: impact of season, location and environment. Polar Biology, DOI:10.1007/s00300-015-1725-9
  11. Zarubin, M., Genin, A., Falk-Petersen, S., Wold, A., Farstey, V. 2014. Intraspecific differences in lipid content of Calanoid copepods across fine-scale depth ranges within the photic layer. Plos One 9(3). DOI:10.1371/journal.pone.0092935
  12. Blachowiak-Samolyk, K., Anna-Maria, K., Falk-Petersen, S., Wold, A., Hegseth, E.N., Wiktor, J. 2014. Winter Tales: The dark side of planktonic life. Polar Biology 38(1):23-36. DOI:10.1007/s00300-014-1597-4
  13. Daase, M., Falk-Petersen, S., Varpe, Ø., Darnis, G., Søreide, J.E., Wold, A., Leu, E., Berge, J., Philippe, B., Fortier, L. 2013. Timing of reproductive events in the marine copepod Calanus glacialis: a pan-Arctic perspective. Canadian Journal of Fisheries and Aquatic Sciences 70(6): 871–884. DOI:10.1139/cjfas-2012-0401
  14. Berge, J., Varpe, Ø., Moline, M.A., Wold, A., Renaud, P.E., Daase, M., Falk-Petersen, S. 2012. Retention of ice-associated amphipods: possible consequences for an ice-free Arctic Ocean. Biology Letters 8(6): 1012–1015.
  15. Wold, A., Darnis, G., Soreide, J.E., Leu, E., Philippe, B., Fortier, L., Poulin, M., Kattner, G., Graeve, M., Falk-Petersen, S. 2011. Life strategy and diet of Calanus glacialis during the winter-spring transition in Amundsen Gulf, south-eastern Beaufort Sea. Polar Biology 34(12): 1929–1946. DOI:10.1007/s00300-011-1062-6
  16. Rabindranath, A., Daase, M., Falk-Petersen, S., Wold, A., Wallace, M.I., Berge, J., Brierley, A.S. 2011. Seasonal and diel vertical migration of zooplankton in the High Arctic during autumn midnight sun of 2008. Marine Biodiversity 41(3): 365–382. DOI:10.1007/s12526-010-0067-7
  17. Wold, A., Jæger, I., Hop, H., Gabrielsen, G.W., Falk-Petersen, S. 2011. Arctic seabird food chains explored by fatty acid composition and stable isotopes in Kongsfjorden, Svalbard. Polar Biology 34(8): 1147–1155.
  18. Falk-Petersen, S., Leu, E., Berge, J., Kwasniewski, S., Nygard, H., Rostad, A., Keskinen, E., Thormar, J., Quillfeldt, C.H.v., Wold, A., Gulliksen, B. 2008. Vertical migration in high Arctic waters during autumn 2004. Deep Sea Research Part II: Topical Studies in Oceanography 55(20-21): 2275–2284.
  19. Falk-Petersen, S., Haug, T., Hop, H., Nilssen, K.T., Wold, A. 2008. Transfer of lipids from plankton to blubber of harp and hooded seals off East Greenland. Deep Sea Research Part II: Topical Studies in Oceanography 56(21-22): 2080–2086.
  20. Wold, A., Leu, E., Walkusz, W., Falk-Petersen, S. 2007. Lipids in copepodite stages of Calanus glacialis. Polar Biology 30(5): 655–658.
  21. Cottier, F.R., Tarling, G.A., Wold, A., Falk-Petersen, S. 2006. Unsynchronized and synchronized vertical migration of zooplankton in a high arctic fjord. Limnology and Oceanography 51(6): 2586–2599.
  22. Willis, K., Cottier, F., Kwasniewski, S., Wold, A., Falk-Petersen, S. 2006. The influence of advection on zooplankton community composition in an Arctic fjord (Kongsfjorden, Svalbard). Journal of Marine Systems 61(1-2): 39–54.
  23. Falk-Petersen, S., Haug, T., Nilssen, K.T., Wold, A., Dahl, T.M. 2004. Lipids and trophic linkages in harp seal (Phoca groenlandica) from the eastern Barents Sea. Polar Research 23(1): 43–50. DOI:10.1111/j.1751-8369.2004.tb00128.x
  24. Wold, A., Norrbin, F. 2004. Vertical migration as a response to UVR stress in Calanus finmarchicus females and nauplii. Polar Research 23(1): 27–34.


Photo of Magdalena Wutkowska

Magdalena Wutkowska

PhD candidate

PhD project title: Arctic microbial diversity and functional importance in space and time.…Read More
UNIS – The University Centre in Svalbard Work address Department for Arctic Biology UNIS – The University Centre in Svalbard PO box 156 Longyearbyen 9171 Norway

PhD project title:

Arctic microbial diversity and functional importance in space and time.

PhD project description:

Exploring seasonal changes in functionality of microbial eukaryotes through comparative metatranscriptomics. Last years brought in a lot of understanding of marine microbial communities in High-Arctic marine habitats in Svalbard. The aim of this project is to build up additional layer of understanding by exploring functionality of microbial communities in a perspective of seasonal changes. To focus on the function of whole communities and its correlation with environmental parameters both meta-transcriptomics and comparative meta-transcriptomics approach will be implemented. This could reveal new insights into the biogeochemical significance of 0.45-10 um sized components of marine microbial eukaryotes. Previous analyses have already answered the questions of  ‘who is there?’ and ‘who is active?’, and in the current project it would be possible to assess what physiological processes occur at certain time points and who is responsible for this contribution. This project would involve development of a bioinformatics pipeline for the purpose of analyzing metatranscriptomics data sets. Specific questions to be answered are:

– How does the function of 0.45-10 um fraction of marine microbial communities assemblages change throughout a year?
– What are the main functions of 0.45-10 um fraction of marine microbial communities?
– Are there any correlations between physiological pathways and environmental factors?
– What is the biogeochemical importance and significance of picoeukaryotes in High Arctic waters

Start date: 11.01.2016
Planned submission date: 10.01.2020


Pernille B. Eidesen
Tove M. Gabrielsen
Dorothee Ehrich

Photo of Kasia Zamelczyk

Kasia Zamelczyk


Research interests:
  • Marine microfossils (planktic foraminifera and diatom ecology)
  • Ocean acidification
  • Dissolution of carbonates and carbon cycle
  • Paleoclimate and palaeoceanography reconstruction.
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UiT The Arctic University of NorwayGeology Work address Centre for Arctic Gas Hydrate, Environment and Climate Department of Geology UiT The Arctic University of Norway Tromsø 9037 Norway Work Phone: +47 77 64 47 73 Website: CAGE – Centre for Arctic Gas Hydrate, Environment and Climate

Research interests:

  • Marine microfossils (planktic foraminifera and diatom ecology)
  • Ocean acidification
  • Dissolution of carbonates and carbon cycle
  • Paleoclimate and palaeoceanography reconstruction.

Running national and international projects:

  • Effects of ocean chemistry changes on planktonic foraminifera in the Fram Strait: Ocean Acidification from natural to anthropogenic changes

Completed national and international projects:

  • SciencePub project: Arctic Natural Climate and Environmental Changes and Human Adaptation: From Science to Public Awareness

Relevant scientific publications:

  1. Zamelczyk, K., Husum, K., Rasmussen, T.L., Godtliebsen, T.L., Hald, M., 2014. Surface water conditions and calcium carbonate preservation in the Fram Strait during marine isotope stage 2, 28.8–15.4 kyr, Paleoceanography 29, doi: 10.1002/2012PA002448.
  2. Zamelczyk, K., Rasmussen, T.L., Husum, K., Hald, M., 2013. Marine calcium carbonate preservation vs. climate change over the last two millennia in the Fram Strait: Implications for planktic foraminiferal paleostudies. Marine Micropaleontology, doi: 10.1016/j.marmicro.2012.10.001
  3. Zamelczyk, K., Rasmussen, T.L, Husum, K., Haflidason, H., de Vernal, A., Ravna, E.K., Hald, M., Hillaire-Marcel, C., 2012. Paleoceanographic changes and calcium carbonate dissolution in the central Fram Strait during the last 20 ka. Quaternary Research, 78, 405-416, doi:10.1016/j.yqres.2012.07.006
  4. Werner, K., Spielhagen, R.F, Bauch, D., Hass, H.Ch., Kandiano, E.,  Zamelczyk, K., 2011. Atlantic Water advection to the eastern Fram Strait-multiproxy evidence for late Holocene variability. Palaeogeography Palaeoclimatology Palaeoecology 308(3-4), 264-276, doi:10.1016/j.palaeo.2011.05.030.
  5. Spielhagen, R.F, Werner, K., Aagaard-Sørensen, S., Zamelczyk, K., Kandiano, E., Budeus, G., Husum, K., Marchitto, T., Hald, M., 2011. Enhanced modern heat transfer to the Arctic by warm Atlantic Water. Science 331, 450, doi: 10.1126/science.1197397.

Popular science and outreach:

  • Zamelczyk, K., Arctic climate variations in geological perspective- oral presentations during     the “Researches Night” on board MS North Star, Hurtigrutene, 24-25.09.2010
  • Jessen, Simon Pind; Sørensen, Steffen Aagaard; Zamelczyk, Katarzyna. Oceanography, Palaeoceanography and Climate. Part of the Teacher’s Guide to the exhibition TRACK – On the trail of past climate. [Artistic and museum presentation] TRACK – The track of past climate. Science Center in Trondheim, Trondheim, and 6 other cities 2009-2013. 2009
  • Jessen, Simon, Pind , Aagaard-Sørensen, S., Zamelczyk, K. Exhibitions and presentations of current research state during “Science days”, Tromsø, Norway, 2008
  • Jessen, Simon, Pind, Aagaard-Sørensen, S., Zamelczyk, K.,  Expedition exhibition. Traveling plaeogeological exhibition for middle school pupils. Nordnorsk Vitensenteret (Norwegian Science Center of Northern Norway, Tromsø), 2008-2011


Photo of Irina Zhulay

Irina Zhulay

PhD candidate

PhD Project title: Arctic Deep-sea benthos: Biodiversity, food web, and biological and functional traits.…Read More
UiT The Arctic University of NorwayArctic and Marine Biology Work address Department of Arctic and Marine Biology Faculty of Biosciences, Fisheries and Economics UiT The Arctic University of Norway Tromsø 9034 Norway Work Phone: +47 776 44492

PhD Project title:

Arctic Deep-sea benthos: Biodiversity, food web, and biological and functional traits.

PhD Project description:

My PhD project focuses on the description of deep sea faunal characteristics in the Arctic. The Arctic deep sea is currently one of the least explored regions. It is also a region that is in a rapid state of transition, which is linked to the decline in ice coverage. This makes description of Arctic deep-sea ecosystem an urgent need. The overall aim of my thesis is to improve current knowledge of the deep-sea fauna distribution and characterize deep-sea communities in terms of food web structure and functional trait composition. The first thesis chapter will explain food web structure in the central Arctic, specifically the Canada Basin including a comparison of Amerasian and Eurasian Basin zooplankton. Stable isotope analysis will is used to examine deep-sea food webs. The second chapter will deal with a description of biological and functional traits of select Arctic deep-sea benthos. The main approach for this chapter will be the Biological Trait Analysis (BTA), which uses biological traits such as a series of life history, morphological and behavioral characteristics present in a community that are directly related to mechanisms structuring the ecosystems, and, thereby, can be used to describe ecological functioning of ecosystems. The third chapter will improve knowledge on epifauna community structure in the heterogeneous Chukchi Borderland based on ROV imagery and bottom trawls.

Start date: Nov 2015
End date: Autumn 2019


Bodil Bluhm (UiT, Tromsø, Norway)
Paul Renaud (Akvaplan-niva, Tromsø, Norway)

Relevant scientific publications:

  1. Zhulay I, Reiss K, Reiss H (2015) Effects of aquaculture fallowing on the recovery of macrofauna communities. Marine Pollution Bulletin 97 (1-2): 381 – 390 Link to article.
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