Kit Kovacs » Members

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Kit Kovacs

Senior Scientist Norwegian Polar InstituteResearch department: Biodiversity
Work address Norwegian Polar Institute Fram Centre Tromsø 9296 Norway Work Phone: +47 77 75 05 26
Photo of Kit Kovacs

Research interests:

  • Marine ecology
  • Arctic marine biology
  • Top predators
  • Polar ecology
  • Marine mammal biology
  • Mammalian evolution.

Running national and international projects:

  • ICE whales (national)
  • MEOP (international)
  • CAFF bearded seal ecology

Relevant scientific publications:

  1. Hansen, M., Villanger, G.D., Bechshoft, T., Levin, M., Routti, H., Kovacs, K.M., Lydersen, C. 2017. Circulating thyroid hormones and associated metabolites in white whales ( Delphinapterus leucas ) determined using isotope-dilution mass spectrometry. Environmental Research 156: 128–131.
    DOI:10.1016/j.envres.2017.03.027
  2. Descamps, S., Aars, J., Fuglei, E., Kovacs, K.M., Lydersen, C., Pavlova, O., Pedersen, Å.Ø., Ravolainen, V., Strøm, H. 2017. Climate change impacts on wildlife in a High Arctic archipelago – Svalbard, Norway. Global Change Biology 23.
    DOI:10.1111/gcb.13381
  3. Vacquié-Garcia, J., Lydersen, C., Marques, T.A., Aars, J., Ahonen, H., Skern-Mauritzen, M., Øien, N., Kovacs, K.M. 2017. Late summer distribution and abundance of ice-associated whales in the Norwegian High Arctic. Endangered Species Research 32.
    DOI:10.3354/esr00791
  4. Grønnestad, R., Villanger, G.D., Polder, A., Kovacs, K.M., Lydersen, C., Jenssen, B.M., Borgå, K. 2017. Maternal transfer of perfluoroalkyl substances in hooded seals. Environmental Toxicology and Chemistry 36: 763–770.
    DOI:10.1002/etc.3623
  5. Scheffers, B.R., De Meester, L., Bridge, T.C.L., Hoffmann, A.A., Pandolfi, J.M., Corlett, R.T., Butchart, S.H.M., Pearce-Kelly, P., Kovacs, K.M., Dudgeon, D., Pacifici, M., Rondinini, C., Foden, W.B., Martin, T.G., Mora, C., Bickford, D., Watson, J.E.M. 2016. The broad footprint of climate change from genes to biomes to people. Science 354.
    DOI:10.1126/science.aaf7671
  6. Tarroux, A., Lowther, A.D., Lydersen, C., Kovacs, K.M. 2016. Temporal shift in the isotopic niche of female Antarctic fur seals from Bouvetøya. Polar Research 35.
    DOI:10.3402/polar.v35.31335
  7. Nyhus, E.S., Lindqvist, C., Kovacs; K.M., Lydersen, L., Wiig, Ø, and Bachmann, L. 201x. Mitogenomics of contemporary Spitsbergen stock bowhead whales (Balaena mysticetus). Mitochondrial DNA Part B, 1:1, 898-900, doi:10.1080/23802359.2016.1258345.
  8. Steeves, H. N., McMeans, B., Field, C., Stewart, C., Arts, M. T., Fisk, A. T., Lydersen, C., Kovacs, K. M. and MacNeil, M. A. 2016. Non-parametric analysis of the spatio-temporal variability in the fatty-acid profiles among Greenland sharks. J. Mar. Biol. Assoc. UK. – in press October 2016.
  9. Kovacs, K.M., Lemons, R. and Lydersen, C. 2016. Walruses in a time of climate change.  Bull. Am. Met. Soc. S97 (No 8): 136-137.
  10. Lowther, A.D., Lydersen, C., Kovacs, K.M. 2016. The seasonal evolution of shelf water masses around Bouvetøya, a sub-Antarctic island in the mid-Atlantic sector of the Southern Ocean, determined from an instrumented southern elephant seal. Polar Research 35.
    DOI:10.3402/polar.v35.28278
  11. Lydersen, C., Fisk, A.T., Kovacs, K.M. 2016. A review of Greenland shark (Somniosus microcephalus) studies in the Kongsfjorden area, Svalbard Norway. Polar Biology 39.
    DOI:10.1007/s00300-016-1949-3
  12. Hamilton, C., Lydersen, C., Ims, R.A., Kovacs, K.M. 2016. Coastal habitat use by ringed seals (Pusa hispida) following a regional sea-ice collapse – the importance of glacial refugia in a changing Arctic. Marine Ecology Progress Series 545: 261–277.
    DOI:10.3354/meps11598
  13. Routti, H., Gabrielsen, G.W., Herzke, D., Kovacs, K.M., Lydersen, C. 2016. Spatial and temporal trends in perfluoroalkyl substances (PFASs) in ringed seals (Pusa hispida) from Svalbard. Environmental Pollution 214.
    DOI:10.1016/j.envpol.2016.04.016
  14. Lindqvist, C., Roy, T., Lydersen, C., Kovacs, K.M., Aars, J., Wiig, Ø., Bachmann, L. 2016. Genetic diversity of historical Atlantic walruses (Odobenus rosmarus rosmarus) from Bjørnøya and Håøya (Tusenøyane), Svalbard, Norway. BMC Research Notes 9: 1–8.
    DOI:10.1186/s13104-016-1907-8
  15. Nyhus, E.S., Lindqvist, C., Kovacs, K., Lydersen, C., Wiig, Ø., Bachmann, L. 2016. Mitogenomes of contemporary Spitsbergen stock bowhead whales (Balaena mysticetus). Mitochondrial DNA Part B 1.
    DOI:10.1080/23802359.2016.1258345
  16. Niemandt, C., Kovacs, K.M., Lydersen, C., Dyer, B.M., Isaksen, K., Hofmeyr, G.J.G., Mehlum, F., Bruyn, P.J.N.d. 2016. Chinstrap and macaroni penguin diet and demography at Nyrøysa, Bouvetøya. Antarctic Science 28: 91–100.
    DOI:10.1017/S0954102015000504
  17. Kovacs, K.M., Lemons, P., Lydersen, C. 2016. Walruses in a time of climate change. Bulletin America Meterological Society 136-137 pp.
  18. Hallanger, I.G., Sagerup, K., Evenset, A., Kovacs, K.M., Leonards, P., Fuglei, E., Routti, H., Aars, J., Strøm, H., Lydersen, C., Gabrielsen, G.W. 2015. Organophosphorous flame retardants in biota from Svalbard, Norway. Marine Pollution Bulletin 101(1): 442–447.
    DOI:10.1016/j.marpolbul.2015.09.049
  19. Lowther, A.D., Lydersen, C., Kovacs, K.M. 2015. A sum greater than its parts: merging multi-predator tracking studies to increase ecological understanding. Ecosphere 6(12): 1–13.
    DOI:10.1890/ES15-00293.1
  20. Hamilton, C., Lydersen, C., Ims, R.A., Kovacs, K.M. 2015. Predictions replaced by facts: a keystone species’ behavioural responses to declining Arctic sea-ice. Biology Letters 11(11): 20150803. 6 pp.
    DOI:10.1098/rsbl.2015.0803
  21. Lowther, A.D., Ahonen, H., Hofmeyr, G., Oosthuizen, W.C., De Bruyn, P.J.N., Lydersen, C., Kovacs, K.M. 2015. Reliability of VHF telemetry data for measuring attendance patterns of marine predators: a comparison with time depth recorder data. Marine Ecology Progress Series 538: 249–256.
    DOI:10.3354/meps11504
  22. Blanchet, M.-A., Lydersen, C., Ims, R.A., Kovacs, K.M. 2015. Seasonal, oceanographic and atmospheric drivers of diving behaviour in a temperate seal species living in the high Arctic. Plos One 10(7): 1–28.
    DOI:10.1371/journal.pone.0132686
  23. Lowther, A.D., Lydersen, C., Fedak, M.A., Lovell, P., Kovacs, K.M. 2015. The Argos-CLS Kalman Filter: Error Structures and State-Space Modelling Relative to Fastloc GPS Data. Plos One 10(4): 1–16.
    DOI:10.1371/journal.pone.0124754
  24. Lowther, A.D., Kovacs, K.M., Griffiths, D., Lydersen, C. 2015. Identification of motivational state in adult male Atlantic walruses inferred from changes in movement and diving behavior. Marine Mammal Science 31(4): 1291–1313.
    DOI:10.1111/mms.12224
  25. Laidre, K.L., Stern, H., Kovacs, K.M., Lowry, L., Moore, S.E., Regehr, E.V., Ferguson, S.H., Wiig, Ø., Boveng, P., Angliss, R.P., Born, E.W., Litovka, D., Quakenbush, L., Lydersen, C., Vongraven, D. and Ugarte, F. 2015. Arctic marine mammal population status, sea ice habitat loss, and conservation recommendations for the 21st century. Conservation Biology 29 (3): 724–737. DOI: 10.1111/cobi.12474.
  26. Pacifici, M. Foden, W.B., Visconti, P., Watson, J.E.M., Butchart, S.H.M., Kovacs, K.M., Scheffers, B.R., Hole, D.G., Martin, T.G., Akçakaya, H.R., Corlett, R.T., Huntley, B., Bickford, D., Carr, J.A., Hoffmann, A.A., Midgley, G.F., Pearce-Kelly, P., Pearson, R.G., Williams, S.E., Willis, S.G., Young, B. and Rondinini, C. 2015. Assessing species vulnerability to climate change. Nature Climate Change 5: 215-224. doi: 10.1038.
  27. . Hamilton, C.D., Kovacs, K.M. and Lydersen, C. 2015. Year-round haul-out behaviour of walruses (Odobenus rosmarus) in the Northern Barents Sea. MEPS 519: 251-263, doi: 10.3354/meps11089.
  28. Falk-Petersen, S., Pavlov, V., Berge, J., Cottier, F., Kovacs, K.M. and Lydersen, C. 2015. At the rainbow’s end – productivity hotspots due to upwelling along Arctic shelves. Polar Biology 38: 5-11, doi: 10.1007/s00300-014-1482-1.
  29. Grist, J.P., Josey, S.A., Boehme, L., Meredith, M.P., Laidre, K.L., Heide-Jørgensen, M.P., Kovacs, K.M. and Lydersen, C., Davidson, Stenson, G.B., Hammill, M.O., Marsh, R. and Coward, A.C. 2014. How, when and where does the warmest Atlantic Water reach the Greenland shelf? Geophysical Research Letters 41, 8530–8537, doi:10.1002/2014GL062051.
  30. Tverberg, V., Nøst, O.A., Lydersen, C. and Kovacs, K.M. 2014. Winter sea ice melting in the Atlantic Water subduction area, Svalbard Norway. Journal of Geophysical Research Oceans, 119, 5945-5967. doi:10.1002/2014JC010013.
  31. Routti, H., Lydersen, C., Hanssen, L. and Kovacs, K.M. 2014. Current levels of contaminants in the world’s northernmost harbour seals (Phoca vitulina). Marine Pollution 87: 140-146.
  32. Isachsen, P.E., Sørlie, S.R., Mauritzen, C., Lydersen. C., Dodd, P. and Kovacs, K.M. 2014. Upper-ocean hydrography of the Nordic Seas during the International Polar Year (2007–2008) as observed by instrumented seals and Argo floats. Deep-Sea Research I 93: 41-59, doi: 10.1016/j.dsr.2014.06.012.
  33. Blanchet, M.A., Lydersen, C., Ims, R.A., Lowther, A.D. and Kovacs, K.M. 2014. Harbour seal (Phoca vitulina) movement patterns in the High Arctic archipelago of Svalbard, Norway. Aquatic Biology21: 167-181.
  34. Meier, W.N., Key, J.R., Kovacs, K.M., Michel, C., Hovelsrud, G.K., van Oort, B.E.H., Haas, C., Granskog, M.A., Gerland, S., Perovich, D.J., Makshtas, A. and Reist, J.D. 2014. Arctic sea ice in transformation: A review of recent observed changes and impacts on 1 biology and human activity. Reviews of Geophysics 51: 185-217.
  35. Kovacs, K.M., Aars, J. and Lydersen, C. 2014. Walruses recovering after 60+ years of protection at Svalbard, Norway. Polar Research 33, 26034, doi.org/10.3402/polar.v.33.26034.
  36. Lowther, A.D., Lydersen, C., Biuw, M., Nico de Bruyn, P.J., Hofmeyr, G.J.G. & Kovacs, K.M. 2014. Post-breeding at-sea movement of three central place foragers in relation to submesoscale fronts in the Southern Ocean around Bouvetøya. Antarctic Science 26: 533-544.
  37. Zhou, Q., Hattermann, T., Nøst, O.A., Biuw, M., Kovacs, K.M. and Lydersen, C. 2014. Wind-driven spreading of fresh surface water beneath ice shelves in the Eastern Weddell Sea. Journal of Geophysical Research – Oceans, 119, 3818–3833, doi:10.1002/2013JC009556.
  38. Hamilton, C.D., Lydersen, C., Ims, R.A. and Kovacs, K.M. 2014. Haul-out behaviour of the world’s northernmost population of harbour seals (Phoca vitulina) throughout the year. PLoS ONE 9(1): e86055, doi:10.1371/journal.pone.0086055
  39. Lu, Z., Fisk, A.T., Kovacs, K.M., Lydersen, C., McKinney, M.A., Tomy, G.T., Rosenburg, B., McMeans, B.C., Muir, D.C.G. and Wong, C.S. 2014. Temporal and spatial variation in polychlorinated biphenyl chiral signatures of the Greenland shark (Somniosus microcephalus) and its arctic marine food web. Environmental Pollution 186: 216-225.
  40. Reeves, R.R., Ewins, P.J., Agbayani, S., Heidi-Jorgensen, M.P., Kovacs, K.M., Lydersen, C., Suydam, R., Elliott, W., Polet, G., van Dijk, Y. and Blijleven, R. 2014. Distribution of endemic cetaceans in relation to hydrocarbon development and commercial shipping in a warming Arctic. Marine Policy 44: 375-389.
  41. Lydersen, C., Assmy, P., Falk-Petersen, S., Kohler J., Kovacs, K.M., Reigstad, M., Steen, H., Strøm, H., Sundfjord, A., Varpe, Ø., Walczowski, W., Weslawski, J.M. and Zajaczkowski, M. 2014. The importance of tidewater glaciers for marine mammals and seabirds in Svalbard, Norway. Journal of Marine Systems 129: 452-471.
  42. Blanchet, M.A., Lydersen, C., Biuw, M., Hofmeyr, G., Krafft, B., and Kovacs, K.M. 2014. Instrumentation and handling effects on Antarctic fur seals (Arctocephalus gazella) behaviour. Polar Research 33, 21630, http://dx.doi.org/10.3402/polar.v33.21630.
  43. Roquet, F., Wunsch, C., Forget, C., Heimbach, P., Guinet, C., Reverdin, G., Charrassin, J.-B., Bailleul, F., Costa, D., Huckstadt, L., Goetz, K., Kovacs, K.M., Lydersen, C., Nøst, O.A., Bornemann, H., Ploetz, J., Bester, M.N., McIntyre,T., Muelbert, M.C., Hindell, M., Williams, G., Field, I., Harcourt, R., Nicholls, K., Chafik, L., Biuw, M., Boehme, L. and Fedak, M. 2013. Hydrographic data collected by seals significantly reduce the observational gap in the Southern Ocean. Geophysical Research Letters 40, 6176–6180, doi:10.1002/2013GL058304.
  44. Merkel, B., Lydersen, C., Yoccoz, N.G. and Kovacs, K.M. 2013. The worlds’s northernmost harbour seal population – How many are there? PLoS ONE 8(7): e67576. doi:10.1371/journal.pone.0067576.
  45. Villanger, G.D., Gabrielsen, K.M., Kovacs, K.M., Lydersen, C., Lie, E., Karimi, M., Sørmo, E.G., Jenssen, B.M. 2013. Effects of complex organohalogen contaminant mixtures on thyroid homeostasis in hooded seal (Cystophora cristata) mother-pup pairs. Chemosphere 92(7): 828–842.
    DOI:10.1016/j.chemosphere.2013.04.036
  46. Roth, S. J., Tischer, B. K., Kovacs, K.M., Lydersen, C., Osterrieder, N. and Tryland, M. 2013. Phocine herpesvirus 1 (PhHV-1) in harbor seals from Svalbard, Norway. Vet. Microbiol. (PhHV-1) in harbor seals from Svalbard, Norway. Vet. Microbiol., http://dx.doi.org/10.1016/j.vetmic.2013.03.008.
  47. Lydersen, C., Øien, N., Mikkelsen, B., Bober, S., Fisher, D. and Kovacs, K.M. 2013. A white humpback whale (Megaptera novaeangliae) in the Atlantic Ocean, Svalbard, Norway, August 2012. Polar Research 2013, 32, 19739, http://dx.doi.org/10.3402/polar.v32i0.19739.
  48. McMeans, B.C., Lydersen, C., Fish, A., Arts, M., Hop, H., Falk-Petersen, S. and Kovacs, K.M. 2013. The role of Greenland sharks in an Arctic ecosystem – assessed via fatty acids and stable isotopes. Marine Biology 160: 1223-1238. doi.10.1007/s00227-013-2174-z.
  49. Iversen, M., Aars, J., Haug, T., Alsos, I., Lydersen, C., Bachmann, L. and Kovacs, K.M. 2013. The diet of polar bears (Ursus maritimus) from Svalbard, Norway, inferred from scat analysis. Polar Biology 36: 561-571. Doi. 10.1007/s00300 012 12842.
  50. Lydersen, C. and Kovacs, K.M. 2013. Walrus Odobenus rosmarus research in Svalbard, Norway, 2000-2010. NAMMCO Sci. Publ. in press
  51. Castellote, M., Leeney, R.H., O’Corry Crowe, G., Lauhakangas, R., Kovacs, K.M., Lucey, W., Krasnova, V., Lydersen, C., Stafford, K.M. and Belikov, R. 2013. Monitoring white whales (Delphinapterus leucas) with echolocation loggers. Polar Biology 36: 493-509. doi 10.10071500300 012 12762.
  52. Blanchet, M.A. Biuw, M., Hofmeyr, G.J.G., Nico de Bruyn, P.J., Lydersen, C., and Kovacs, K.M. 2013. At-sea behaviour of three krill predators breeding at Bouvetøya – Antarctic fur seals, macaroni penguins and chinstrap penguins. Marine Ecology Progress Series 477: 285-302.
  53. Molde, K., Ciesielski, T., Fisk, A. T., Lydersen, C., Kovacs, K.M., Sørmo, E. G. and Jenssen, B. M. 2013. Associations between vitamins A and E and legacy POP levels in highly contaminated Greenland sharks (Somniosus microcephalus). Science of the Total Environment 442: 445-454.
  54. Hunt, G.L. Jr., Blanchard, A.L., Boveng, P., Dalpadado, P., Drinkwater, K., Eisner, L., Hopcroft, R., Kovacs, K.M., Norcross, B., Renaud, P., Reigstad, M., Whitehouse, G.A., Woodgate, R. 2013. The Barents and Chukchi Seas Compared: Why does the Barents support more productive fisheries? Comparative Journal of Marine Systems 109-110: 43-68.
  55. Fisk, A.T., Lydersen, C. and Kovacs, K.M. 2012. Archival pop-off tag tracking of Greenland sharks (Somniosus microcephalus) in the High Arctic waters of Svalbard, Norway. Marine Ecology Progress Series – 468: 255-265. DOI 10.3354/meps09962.
  56. Stafford, K.M., Moore, S.E., Berchok, C., Wiig, Ø., Lydersen, C., Hansen, E. and Kovacs, K.M. 2012. Spitsbergen’s endangered bowhead whales sing through the polar night. Endangered species 18: 95-103.
  57. Harvey, V., Hammill, M.O., Swain, D.P., Breed, G.A., Lydersen, C. and Kovacs, K.M. 2012. Winter foraging by a top predator, the grey seal Halichoerus grypus, in relation to the distribution of prey. Marine Ecology Progress Series 462: 273-286.
  58. Lydersen, C., Chernook, V.I., Glazov, D.M., Trukhanova, I.S. and Kovacs, K.M. 2012. Aerial survey of Atlantic walruses (Odobenus rosmarus rosmarus) in the Pechora Sea, August 2011. Polar Biology 35: 1555-1562.
  59. Hindell, M.A., Lydersen, C., Hop, H. and Kovacs, K.M. 2012. Pre-partum diet of adult female bearded seals in years of contrasting ice conditions. PLosONE 7(5): e38307.
  60. Watanabe, Y.Y., Lydersen, C., Fisk, A.T. and Kovacs, K.M. 2012. The slowest fish: swim speed and tail-beat frequency of Greenland sharks. Journal of Experimental Marine Biology and Ecology 426-427: 5-11.
  61. Leclerc, L.M.E., Lydersen, L., Haug, T., Bachmann, L., Fisk, A.T. and Kovacs, K.M. 2012. A missing puzzle piece in Arctic food web puzzle? Stomach contents of Greenland Sharks sampled off Svalbard, Norway. Polar Biology 35: 1197-1208. doi10.1007/s00300-012-1166-7
  62. Tryland, M., Nymo, I.H., Nielsen, O., Nordøy, E.S., Kovacs, K.M., Krafft, B.A., Thoresen, S.I., Åsbakk, K., Osterrieder, K., Roth, S.J., Lydersen, C., Godfroid, J., and Blix, A.S. 2012. Serum chemistry and antibodies against pathogens in Antarctic fur seals, Weddell seals, crabeater seals and Ross seals. Journal of Wildlife Diseases 48(3): 632-645.
  63. MacNeil, M.A., McMeans, B.C., Hussey, N., Vecsei, P., Svavarsson, J., Kovacs, K.M., Lydersen, C., Treble, M., Skomal, G.B., Ramsey, M. and Fisk, A.T. 2012. Biology of the Greenland shark Somniosus microcephalus. Journal of Fish Biology 80: 991-1018.
  64. Gilg, O., Kovacs, K.M., Aars, J., Fort, J., Gauthier, G., Gramillet, D., Ims, R.A., Meltofte, H., Moreau, J., Post, E., Schmidt, N.M., Yannic, G. and Bollache, L. 2012. Climate change and the ecology and evolution of Arctic vertebrates. Annals of the New York Academy of Science – 1249: 166-190. Doi.10.1111/j.1749-6632 2011.0641.x
  65. Freitas, C., Kovacs, K.M., Andersen, M., Aars, J., Sandven, S., Skern-Mauritzen, Pavlova, O. and Lydersen, C. 2012. Importance of fast ice and glacier fronts for female polar bears and their cubs during spring in Svalbard, Norway. Marine Ecology Progress Series 447: 289-304.
  66. Kovacs, K.M., Aguilar, A., Aurioles, D., Burkanov, V., Campagna, C., Gales, N., Gelatt, T., Goldsworthy, S., Goodman, S.J., Hofmeyr, G.J.G., Härkönen, T., Lowry, L., Lydersen, C., Schipper, J., Sipilä, T., Southwell, C., Stuart, S., Thompson, D. and Trillmich, F. 2012. Global threats to pinnipeds. Marine Mammal Science 28: 414-436.
  67. Moore, S.E., Stafford, K.M., Melling, H., Berchok, C., Wiig, Ø., Kovacs, K.M. and Lydersen, C., Richter-Menge, J. 2012. Comparing marine mammal acoustic habitats in Atlantic and Pacific sectors of the High Arctic: year-long records from Fram Strait and the Chukchi Plateau. Polar Biology 35: 475-480.
  68. Lydersen, C, Øystein, Ø, Wiig, Bachmann, L., Heide-Jørgensen, M.P., Swift, R. and Kovacs, K.M. 2012. Lost highway not forgotten: Satellite tracking of a bowhead whale (Balaena mysticetus) from the critically endangered Spitsbergen stock. Arctic 65:76-86.
  69. Vincent, W.F., Callaghan, T., Dahl-Jensen, D., Johansson, M., Kovacs, K.M., Michel, C., Prowse, T., Reist, J.D. and Sharp, M. 2011. Ecological implications of changes in the Arctic Cryosphere. AMBIO 40: 87-99. DOI 10.1007/s13280-011-0218-5.
  70. Nøst, O.-A., Biuw, M., Tverberg, V., Lydersen, C., Zhou, Q., Hattermann, T., Smedsrud, L.-H., and Kovacs, K.M. 2011. Eddy overturning of the Antarctic slope front controls glacial melting in the Eastern Weddell Sea. Journal of Geophysical Research 116, C11014, doi:10.1029/2011JC006965.
  71. Gabrielsen, K.M., Villanger, G.D., Lie, E., Karimi, M., Lydersen, C., Kovacs, K.M., Jenssen, B.M. 2011. Levels and patterns of hydroxylated polychlorinated biphenyls (OH-PCBs) and their associations with thyroid hormones in hooded seal (Cystophora cristata) mother-pup pairs. Aquatic Toxicology 105: 482-491.
  72. Villanger, G.D., Lydersen, C., Kovacs, K.M., Lie, E., Skaare, J.U., Jenssen, B.M. 2011. Disruptive effects of persistent organohalogen contaminants on thyroid function in white whales (Delphinapterus leucas) from Svalbard. Science of the Total Environment 409: 2511-2524.
  73. Leclerc, L.M., Lydersen, C., Haug, T., Glover, K.A. and Kovacs, K.M. 2011. Greenland sharks (Somniosus microcephalus) scavenge offal from minke (Balaenoptera acutorostrata) whaling operations in Svalbard (Norway). Polar Research 30, 7342, DOI: 10.3402/polar.v30o0.7342.
  74. Kovacs, K.M., Moore, S., Overland, J.E. and Lydersen, C. 2011. Impacts of changing sea-ice conditions on Arctic marine mammals. Marine Biodiversity 41: 181-194 (DOI 10.1007/S12526-010-0061-0).
  75. Mauritzen, C., Hansen, E., Andersson; M., Berx, B., Beszczynska-Möller, A., Burud, I., Christensen, K.H., Debernard, J., de Steur, L., Dodd, P., Gerland, S., Godøy, Ø., Hansen, B., Hudson, S., Høydalsvik, F., Ingvaldsen, R., Isachsen, P.E., Kasajima, Y., Koszalka, I., Kovacs, K.M., Køltzow, M., LaCasce, J., Lee, C.M., Lavergne T., Lydersen, C., Nicolaus, M., Nilsen, F., Nøst, O.A., Orvik, K.A., Reigstad, M., Schyberg, H., Seuthe, L., Skagseth, Ø., Skarðhamar, J., Skogseth, R., Sperrevik, A., Svensen, C., Søiland, H., Teigen, S.H., Tverberg, V., Wexels Riser, C. 2011. Closing the Loop – Approaches to monitoring the state of the Arctic Mediterranean during the International Polar Year 2007–2008. Progress in Oceanography 90: 62-89.
  76. Andersen, L.W., Kovacs, K.M., Frie, A.K., Rosing-Asvid, A., Hauksson, E. and Lydersen, C. 2011. A population on the edge: genetic diversity and population structure of the world’s northernmost harbor seals (Phoca vitulina). Biological Journal of the Linnean Society 102: 420-439.
  77. Labansen, A.L., Lydersen, C., Levermann, N., Haug, T. and Kovacs, K.M. 2011. Diet of ringed seals (Pusa hispida) from Northeast Greenland. Polar Biology 34: 227-234.
  78. Olsen, M.T., Hirsch-Volny, V., Berube, M., Rew, M.B., Dietz,R., Lydersen, C., Kovacs, K.M., Dodd, R.S. and Palsbøll, P.J. 2011. A simple route to single nucleotide polymorphisms: identification and characterization of 768 SNPs in the ringed seals (Pusa hispida). Molecular Ecology Resources 11 (Suppl. 1): 1-11, doi:10.1111/j.1755-0998.2010.02941.x
  79. Biuw, M. Nøst, Ø.A., Stien, A., Zhou, Q., Lydersen, C. and Kovacs, K.M. 2010. Effects of hydrographic variability on the spatial, seasonal and diel diving patterns of southern elephant seals in the eastern Weddell Sea. PLosOne 5(11): 114 (e13816).
  80. Garde, E., Frie, A.K., Dunshea, G., Hansen, S.H., Kovacs, K.M. and Lydersen, C. 2010. Harp seal ageing techniques – teeth, aspartic acid racemization, and telomere sequence analysis. Journal of Mammalogy 91(6): 1365-1374.
  81. Hoffmann, M., Hilton-Taylor,C., Angulo, A., Böhm, M., Brooks, T.M., Butchart, S.H.M., Carpenter, K.E., Chanson, J., Collen, B., Cox, N.A., Darwall, W.R.T., Dulvy, N.K., Harrison, L.R., Katariya, V., Pollock, C.M., Quader, S., Richman, N.I., Rodrigues, A.S.L., Tognelli, M.F., Vié, J-C., Aguiar, J.M., Allen, D.J., Allen, G.R., Amori, G., Ananjeva, N.B., Andreone, F., Andrew, P., Luz Aquino Ortiz, A., Baillie, J.E.M., Baldi, R., Bell, B.D., Biju, D., Bird, J.P., Black-Decima, P., Blanc, J.J., Bolaños, F., Bolivar, G.W., Burfield, I.J., Burton, J.A., Capper, D.R., Castro, F., Catullo, G., Cavanagh, R.D., Channing, A., Chao, N.L., Chenery, A.M., Chiozza, F., Clausnitzer, V., Collar, N.J., Collett, L.C., Collette, B.B., Cortez Fernandez, C.F., Craig, M.T., Crosby, M.J., Cumberlidge, N., Cuttelod, A., Derocher, A.E., Diesmos, A.C., Donaldson, J.S., Duckworth, J.W., Dutson, G., Dutta, S.K., Emslie, R.H., Farjon, A., Fowler, S., Freyhof, J., Garshelis, D.L., Gerlach, J., Gower, D.J., Grant, T.D., Hammerson, G.A., Harris, R.B., Heaney, L.R., Hedges, S.B., Hero, J-M., Hughes, B., Hussain, S.A., Icochea; J.M., Inger, R.F., Ishii, N., Iskandar, D.T., Jenkins, R.K.B., Kaneko, Y., Kottelat, M., Kovacs, K.M., Kuzmin, S.L., La Marca, E., Lamoreux, J.F., Lau, M.W.N., Lavilla, E.O., Leus, K.L., Lewison, R.L., Lichtenstein, G., Livingstone, S.R., Lukoschek, V., Mallon, D.P., McGowan, P.J.K., McIvor, A., Moehlman, P.D., Molur, S., Muñoz Alonso, A., Musick, J.A., Nowell, K., Nussbaum,R.A., Olech, W., Orlov, N.L., Papenfuss, T.J., Parra-Olea, G., Perrin, W.F., Polidoro, B.A., Pourkazemi, M., Racey, P.A., Ragle, J.S., Ram, M., Rathbun, G., Reynolds, R.P., Rhodin, A.G.J., Richards, S.J., Rodríguez, L.O., Ron, S.R., Rondinini, C., Rylands, A.B., Sadovy de Mitcheson, Y., Sanciangco, J.C., Sanders, K.L., Santos-Barrera, G., Schipper, J., Self-Sullivan, C., Shi, Y., Shoemaker, A., Short, F.T., Sillero-Zubiri, C., Silvano, D.L., Smith, K.G., Smith, A.T., Snoeks, J., Stattersfield, A.J., Symes, A.J., Taber, A.B., Talukdar, B.K., Temple, H.J., Timmins, R., Tobias, J.A., Tsytsulina, K., Tweddle, D., Ubeda, C., Valenti, S.V., van Dijk, P.P., Veiga, L.M., Veloso, A., Wege, D.C., Wilkinson, M., Williamson, E.A., Xie, F., Young, B.E., Akçakaya, R., Bennun, L., Blackburn, T.M., Boitani, L., Dublin, H.T., da Fonseca, G.A.B., Gascon, C., Lacher, T.E., Jr., Mace, G.M., Mainka, S.A., McNeely, J.A., Mittermeier, R.A., McGregor Reid, G., Rodriguez, J.P., Rosenberg, A.A., Samways, M.J., Smart, J., Stein, B.A. and Stuart, S.N. 2010. The Impact of Conservation on the Status of the World’s Vertebrates. Science 330: 1503-1509.
  82. Johansen, C.E., Lydersen, C., Aspholm, P., Haug, T. and Kovacs, K.M. 2010.  Helminth parasites in ringed seals (Pusa hispida) from Svalbard, Norway with special emphasis on nematodes: variation with age, sex and location of host. Journal of Parasitology 96(5): 946-953.
  83. Hofmeyr, G.J.G., Bester, M.N., Kirkman, S.P., Lydersen, C. and Kovacs, K.M.  2010.  Intraspecific differences in the diet of Antarctic fur seals at Nyrøysa, Bouvetøya. Polar Biology 33: 1171-1178.
  84. O’Corry-Crowe, G.M., Lydersen, C., Heide-Jørgensen, M.-P., Hansen, L., Mukhametov, L.M., Dove, O. and Kovacs, K.M.  2010.  Population genetic structure and evolutionary history of North Atlantic beluga whales (Delphinapterus leucas) from West Greenland, Svalbard and the White Sea. Polar Biology 33: 1179-1194.
  85. Wiig, Ø., Bachmann, L., Øien, N., Kovacs, K.M. and Lydersen, C.  2010.  Observations of bowhead whales (Balaena mysticetus) in the Svalbard area 1940 – 2009. Polar Biology 33: 979-984.
  86. Skoglund, E.G., Lydersen, C., Grahl-Nielsen, O., Haug, T. and Kovacs, K.M.  2010.  Fatty acid composition of the blubber and dermis of adult male Atlantic walrus (Odobenus rosmarus rosmarus) in Svalbard, and their potential prey. Marine Biology Research 6: 239-250.
  87. Jensen, S.K., Lydersen, C., Kovacs, K.M., Åsbakk, K. and Aars, J.  2010.  The prevalence of Toxoplasma gondii in polar bears and their marine mammalian prey; evidence for a marine transmission pathway? Polar Biology 33: 599-606.
  88. Biuw, M., Lydersen, C., deBruyn, N., Arriola, A., Hofmeyr, G., Kritzinger, P., Kovacs, K.M.  2010.  Long-range migration of a chinstrap penguin from Bouvetøya to Montagu Island (South Sandwich Islands). Antarctic Science 22(2): 157-162.
  89. Routti, H., Jenssen, B.M., Lydersen, C., Bäckman, C., Arukwe, A., Nyman, M., Kovacs, K.M. and Gabrielsen, G.W.  2010. Hormone, vitamin and contaminant status during the moulting/fasting period in ringed seals (Pusa [Phoca]hispida) from Svalbard. Comparative Biochemistry and Physiology 155 (1): 70-76.
  90. Gales, N., Bowen, W.D., Johnston, D., Kovacs, K.M., Littman, C., Perrin, W., Reynolds, J. and Thompson, P.M.  2009.  Guidelines for the treatment of marine mammals in research. Marine Mammal Science 25(3): 725-736.
  91. Biuw, M.E., Krafft, B.A., Hofmeyr, G., Lydersen, C. and Kovacs, K.M.  2009.  Time budgets and at-sea behaviour of lactating female Antarctic fur seals (Arctocephalus gazella) at Bouvetøya. Marine Ecology Progress Series 385: 271-284.
  92. Kinoshita, M., Ohta, H., Higaki, K., Kojima, Y., Urashima, T., Nakajima, K., Suzzuki, M., Kovacs, K.M., Lydersen, C., Hayakawa, T. and Kakehi, K.  2009.  Structural characterization of multi-branched oligosaccharides from seal milk by combination of off-line high-performance liquid chromatography-matris-assisted laser desorption/ionization time-of-flight mass spectrometry and sequential exoycosidase digestion. Analytical Biochemistry 388: 242-253.
  93. Tryland, M., Lydersen, C., Kovacs, K.M. and Thoresen, S.I.  2009.  Serum chemistry reference values in free-ranging North Atlantic male walruses (Odobenus rosmarus rosmarus) from the Svalbard Archipelago. Veterinary Clinical Pathology 38 (4): 501-506.
  94. Watanabe, Y., Lydersen, C., Sato, K., Naito, Y., Miyazaki, N. and Kovacs, K.M.  2009.  Diving behavior and swimming style of nursing bearded seal pups. Marine Ecology Progress Series 380: 287-294.
  95. Lindqvist, C., Bachmann, L., Andersen, L.W., Born, E.W., Arnason, U., Kovacs, K.M., Lydersen, C., Abramov, A.V. and Wiig, Ø.  2009.  The Laptev Sea walrus Odobenus rosmarus laptevi: an enigma revisited. Zoologica Scripta 38: 113-127.
  96. Freitas, C., Kovacs, K.M., Ims, R.A., Fedak, M.A. and Lydersen, C.  2009.  Deep into the ice: over-wintering and habitat selection in Atlantic walruses. Marine Ecology Progress Series 375: 247-261.
  97. Schipper, J., Chanson, J., Chiozza, F., Cox, N., Hoffmann, M., Katariya, V., Lamoreux, J., Rodrigues, A., Stuart, S.N., Temple, H.J., Baillie, J., Boitani, L., Lacher, T.E., Mittermeier, R.A., Smith, A.T., Absolon, D., Aguiar, J.M., Amori, G., Bakkour, N., Baldi, R., Berridge, R.J., Bielby, J., Black, P.A., Blanc, J.J., Brooks, T.M., Burton, J.A., Butynski, T.M., Catullo, G., Chapman, R., Cokeliss, Z., Collen, B., Conroy, J., Cooke, J.G., da Fonseca, G.A.B., Derocher, A.E., Dublin, H., Duckworth, J.W., Emmons, L., Emslie, R.H., Festa-Bianchet, M., Foster, M., Foster, S., Garshelis, D.L., Gates, C., Gimenez-Dixon, M., Gonzalez, S., Gonzalez-Maya, J.F., Good, T.C., Hammerson, G., Hammond, P.S., Happold, D., Happold, M., Hare, J., Harris, R.B., Hawkins, C.E., Haywood, M., Heaney, L., Hedges, S., Helgen, K.M., Hilton-Taylor, C., Hussain, S.A., Ishii, N., Jefferson, T.A., Jenkins, R.K.B., Johnston, C.H., Keith, M., Kingdon, J., Knox, D., Kovacs, K.M., Langhammer, P., Leus, K., Lewison, R., Lichtenstein, G., Lowry, L.F., Macavoy, Z., Mace, G., Mallon, D.P., Masi, M., McKnight, M.W., Medellín, R., Medici, P., Mills, G., Moehlman, P.D., Molur, S., Mora, A., Nowell, K., Oates, J.F., Olech, W., Oliver, W.R.L., Oprea, M., Patterson, B., Perrin, W.F., Polidoro, B.A., Pollock, C., Powel, A., Protas, Y., Racey, P., Ragle, J., Ramani, P., Rathbun, G., Reeves, R.R., Reilly, S.B., Reynolds, J.E. III, Rondinini, C., Rulli, M., Rylands, A.B., Savini, S., Schank, C.J., Sechrest, W., Self-Sullivan, C., Shoemaker, A., Sillero-Zubiri, C., De Silva, N., Smith, D.E., Srinivasulu, C., Stephenson, P.J., van Strien, N., Talukdar, B.K., Taylor, B.L., Timmins, R., Tirira, D.G., Tognelli, M.F., Tsytsulina, K. Veiga, L.M., Vie1, J.-C., Williamson, L., Wyatt, S.A., Xie, Y. and Young, B.E.  2008.  The biogeography of diversity, threat, and knowledge in the world’s terrestrial and aquatic mammals. Science 322: 225-230.
  98. Freitas, C., Kovacs, K.M., Ims, R.A. and Lydersen, C. 2008. Predicting habitat use by ringed seals (Phoca hispida) in a warming Arctic. Ecological Modelling 217: 19-32.
  99. Freitas, C., Kovacs, K.M., Lydersen, C. and Ims, R.A. 2008. A novel method for quantifying habitat selection and predicting habitat use. Journal of Applied Ecology 45: 1213-1220.
  100. Kovacs, K.M. and Lydersen, C. 2008. Climate change impacts on seals and whales in the North Atlantic Arctic and adjacent shelf seas. Science Progress 91(1): 117-150.
  101. Wolkers, H., Krafft, B., van Bavel, B., Helgason, L.B., Lydersen, C. and Kovacs, K.M. 2008. Biomarker responses and decreasing contaminant levels and in ringed seals (Pusa hispida) from Svalbard, Norway. Journal of Toxicology and Environmental Health 71: 1009-1018.
  102. Strandberg, U., Käkelä, A., Lydersen, C., Kovacs, K.M., Grahl-Nielsen, O., Hyvärinen, H., Käkelä, R. 2008. Stratification, composition and function of marine mammal blubber: The ecology of fatty acids in marine mammals. Physiological and Biochemical Zoology 81(4): 473-485.
  103. Freitas, C., Lydersen, C., Fedak, M.A. and Kovacs, K.M. 2008. A simple new algorithm to filter marine mammal Argos locations. Marine Mammal Science 24(2): 315-325.
  104. Lydersen, C., Aars, J. and Kovacs, K.M. 2008. Estimating the number of walruses in Svalbard from aerial surveys and behavioural data from satellite telemetry. Arctic 61: 119-128.
  105. Marshall, C.D., Kovacs, K.M. and Lydersen, C. 2008. Feeding kinematics, suction and hydraulic jetting capabilities in bearded seals (Erignathus barbatus). Journal of Experimental Biology 211:699-708.
  106. Freitas, C., Kovacs, K.M., Ims, R.A., Fedak, M.A. and Lydersen, C. 2008. Ringed seal post-moulting movement tactics and habitat selection. Oecologia 155: 193-204.
  107. Prestrud, K.W., Åsbakk, K., Fuglei, E., Mørk, T., Stien, A., Ropstad, E., Tryland, M., Gabrielsen, G.W., Lydersen, C., Kovacs, K.M., Loonen, M., Sagerup, K. and Oksanen, A. 2007. Serosurvey for Toxoplasma gondii in arctic foxes and possible sources of infection in the high Arctic of Svalbard. Veterinary Parasitology 150: 6-12.
  108. Labansen, A.L., Lydersen, C., Haug, T. and Kovacs, K.M.  2007. Spring diet of ringed seals (Phoca hispida) from north-western Spitsbergen, Norway. ICES Journal of Marine Science 64: 1246-1256.
  109. Wiig, Ø., Bachmann, L., Janik, V.M., Kovacs, K.M. and Lydersen, C. 2007. Spitsbergen bowhead whales revisited. Marine Mammal Science 23(3): 688-693.
  110. Risch, D., Clark, C.W., Corkeron, P.J., Elepfandt, A., Kovacs, K.M., Lydersen, C., Stirling, I. and Van Parijs, S.M.  2007. Vocalisations of male bearded seals, (Erignathus barbatus): classification and geographical variation. Animal Behaviour 73: 747-762.
  111. Lydersen, C., Martin, T., Gjertz, I. and Kovacs, K.M.  2007. Satellite tracking and diving behaviour of sub-adult narwhals (Monodon monoceros) in Svalbard, Norway. Polar Biology 30: 437-442.
  112. Krafft, B.A., Kovacs, K.M. and Lydersen, C. 2007. Distribution of sex and age groups of ringed seals (Pusa hispida) in the fast-ice breeding habitat. Marine Ecology Progress Series 336: 199-206.
  113. Tryland, M., Krafft, B.A., Lydersen, C., Kovacs, K.M. and Thoresen, S.I. 2006. Serum chemistry values for free-ranging ringed seals (Pusa hispida) in Svalbard. Veterinary Clinical Pathology 35(4): 405-412.
  114. Gwynn, J.P., Brown, J.E., Kovacs, K.M. and Lydersen, C.  2006. The derivation of radionuclide transfer parameters for and dose-rates to an adult ringed seal (Phoca hispida) in an Arctic environment. Journal of Environmental Radioactivity 90: 197-209.
  115. Hofmeyr, G.J., Bester, M., Kirkman, S.P., Lydersen, C. and Kovacs, K.M. 2006. Entanglement of Antarctic fur seals at Bouvetøya, Southern Ocean. Marine Pollution Bulletin 52: 1077-1080.
  116. Sørmo, E.G., Pettersvik Salmer, M., Jenssen, B.M., Hop, H., Bæk, K., Lydersen, C., Falk-Petersen, S., Gabrielsen, G.W., Kovacs, K.M., Lie, E. and Skaare, J.U. 2006. Biomagnification of polybrominated diphenyl ether and hexabromocyclododecane flame retardants in the polar bear food chain in Svalbard, Norway. Environmental Toxicology and Chemistry 25(9): 2502-2511.
  117. Wolkers, H., van Bavel, B., Ericson, I., Skoglund, E., Kovacs, K.M. and Lydersen, C. 2006. Congener-specific accumulation and patterns of chlorinated and brominated contaminants in adult male walruses from Svalbard, Norway: indications for individual-specific prey selection. Science of the Total Environment 370: 70-79.
  118. Davies, C., Kovacs, K.M., Lydersen, C. and VanParijs, S.M.  2006. Development of display behaviour in young captive bearded seals (Erignathus barbatus). Marine Mammal Science 22(4): 952-965.
  119. Krafft, B.A., Lydersen, C. and Kovacs, K.M. 2006. Serum haptoglobin concentrations in ringed seals (Pusa hispida) from Svalbard, Norway. Journal of Veterinary Diseases 42(2): 442-446.
  120. Andersen, M., Gwynn, J.P., Dowdall, M., Kovacs, K.M. and Lydersen, C.  2006.  Radiocaesium (137 Cs) in marine mammals from Svalbard, the Barents Sea and the North Greenland Sea.  Science of the Total Environment 363: 87-94.
  121. Brown, J., Dowdall, M., Gwynn, J.P., Børettzen, P., Selnæs, Ø.G., Kovacs, K.M. and Lydersen, C.  2006. Probabilistic biokinetic modelling of radiocaesium uptake in Arctic seal species: verification of modelled data with empirical observations. Journal of Radiation 88: 289-305.
  122. Tryland, M., Thoresen, S.I., Kovacs, K.M. and Lydersen, C.  2006. Serum chemistry of free-ranging white whales (Delphinapterus leucas) in Svalbard.  Veterinary Clinical Pathology 35(2): 199-203.
  123. Krafft, B.A., Kovacs, K.M., Haug, T. and Lydersen, C.  2006. Growth and population parameters of ringed seals (Pusa hispida) from Svalbard, Norway, 2002 – 2004. ICES Journal of Marine Science, 63(6): 1136-1144.
  124. Andersen, G., Foreid, S, Skaare, J.U., Jenssen, B.M., Lydersen, C. and Kovacs, K.M.  2006.  Levels of toxaphenes in white whales (Delphinapterus leucas) from Svalbard, Norway. Science of the Total Environment. Science of the Total Environment 357: 128-137.
  125. Marshall, C.D., Amin, H., Kovacs, K.M. and Lydersen, C.  2006.  Microstructure and innervation of the mystacial vibrissal follicle-sinus complex in bearded seals, Erignathus barbatus (Pinnipedia: Phocidae). Anatomical Record 288A: 13-25.
  126. Krafft, B.A., Ergon, T., Kovacs, K.M., Andersen, M., Aars, J., Haug, T. and Lydersen, C. 2006.  Abundance of ringed seals (Pusa hispida) in the fjords of Spitsbergen, Svalbard, during the peak moulting period. Marine Mammal Science 22(2): 394-412.
  127. Wolkers, H., Lydersen, C., Kovacs, K.M., Burkow, I. and van Bavel, B. 2006. Accumulation, metabolism, and food-chain transfer of chlorinated and brominated contaminants in subadult white whales (Delphinapterus leucas) and narwhals (Monodon monoceros) from Svalbard, Norway.  Archives of Environmental Contaminant Toxicology 50: 69-78.
  128. Carlens, H., Lydersen, C., Krafft, B.A. and Kovacs, K.M.  2006.  Spring haul-out behavior of ringed seals (Pusaa hispida) in Kongsfjorden, Svalbard. Marine Mammal Science 22(2): 379-393.
  129. Hofmeyr, G.J.G., Krafft, B.A., Kirkman, S.P, Bester, M.N., Lydersen, C. and Kovacs, K.M.  2005.  Population changes of Antarctic fur seals at Nyrøysa, Bouvetøya. Polar Biology 28: 725-731.
  130. Birkeland, A., Kovacs, K.M., Lydersen, C. and Grahl-Nielsen, O.  2005. Transfer of fatty acids from mothers to their calves during lactation in white whales (Delphinapterus leucas). Marine Ecology Progress Series 298: 287-294.
  131. Lydersen, C. and Kovacs, K.M.  2005.  Growth and population parameters of the world’s northernmost harbour seals Phoca vitulina residing in Svalbard, Norway. Polar Biology. 28: 156-163.
  132. Pomponi, M., Gavuzzo, E., Bertonati, C., Derocher, A.E., Lydersen, C., Wiig, Ø. and Kovacs, K.M.  2004.  Hemoglobin, pH and DPG/Chloride shifting. Biochimie 86(12): 927-932.
  133. Grahl-Nielsen, O., Andersen, M., Derocher, A.E., Lydersen, C., Wiig, Ø. and Kovacs, K.M.  2004.  Sampling, data treatment and predictions in investigations on fatty acids in marine mammals. Marine Ecology Progress Series 281: 303-306.
  134. Andersen, S.M., Lydersen, C., Grahl-Nielsen, O. and Kovacs, K.M.  2004.  Autumn diet of harbour seals (Phoca vitulina) at Prins Karls Forland, Svalbard, assessed via scat and fatty-acid analyses. Canadian Journal of Zoology 82(8): 1230-1245.
  135. Lesage, V., Hammill, M.O. and Kovacs, K.M.  2004.  Long distance movements of harbour seals (Phoca vitulina) from a seasonally ice-covered area, the St Lawrence River Estuary, Canada.  Canadian Journal of Zoology 82: 1070-1081.
  136. Urashima, T., Nakamura, T., Nakagawa, D., Noda, M., Arai, I., Saito, T., Lydersen, C. and Kovacs, K.M.  2004.  Characterization of oligosaccharides in milk of bearded seal. Comparative Biochemistry and Physiology 138: 1-18.
  137. Wolkers J., van Bavel, B., Derocher, A.E., Wiig, Ø., Kovacs, K.M., Lydersen, C.  and Lindstrøm, G.  2004.  Congener-specific accumulation and food chain transfer of polybrominated diphenyl ethers in two Arctic food chains. Environmental Science and Technology 38: 1667-1674.
  138. Lydersen, C., Nøst, O., Kovacs, K.M. and Fedak, M.A.  2004.  Temperature data from Norwegian and Russian waters of the northern Barents Sea collected by free-living ringed seals.  Journal of Marine Systems 46: 99-108.
  139. Van Parijs, S.M., Lydersen, C. and Kovacs, K.M.  2004.  Effects of ice cover on the behavioural patterns of aquatic-mating male bearded seals.  Animal Behaviour 68: 89-96.
  140. Wolkers, H., Lydersen, C. and Kovacs, K.M.  2004.  Accumulation and lactational transfer of PCBs and pesticides in harbor seals (Phoca vitulina) from Svalbard, Norway.  Science of the Total Environment 319: 137-146.
  141. Grahl-Nielsen, O., Andersen, M., Derocher, A.E., Lydersen, C., Wiig, Ø. and Kovacs, K.M.  2003.  Fatty acid composition of the adipose tissue of polar bears and of their prey: ringed seals, bearded seals and harp seals.  Marine Ecology Progress Series 265: 275-282.
  142. Reder, S., Lydersen, C., Arnold, W. and Kovacs, K.M.  2003.  Haul-out behavior of High Arcic harbour seals (Phoca vitulina vitulina) in Svalbard, Norway.  Polar Biology 27(1): 6-16.
  143. Urashima, T., Nakamura, T., Yamaguchi, K., Munakata, J., Arai, I., Saito, T., Lydersen, C. and Kovacs, K.M.  2003.  Chemical characterization of the oligosaccharides in high arctic harbour seal (Phoca vitulina) milk.  Comparative Biochemistry and Physiology Part A 135: 549-563.
  144. Van Parijs, S.M., Corkeron, P.J., Harvey, J., Hayes, S., Mellinger, D., Rouget, P., Thompson, P.M., Wahlberg, M. and K.M. Kovacs.  2003.  Patterns in vocalizations of male harbor seals. Journal of the Acoustic Society of America 113(6): 3403-3410.
  145. Van Parijs, S.M., Lydersen, C. and Kovacs, K.M.  2003.  Vocalizations and movements suggest alternate mating tactics in male bearded seals.  Animal Behaviour 65: 273-283.
  146. Van Parijs, S.M., Lydersen, C. and Kovacs, K.M.  2003.  Sound produced by individual white whales, Delphinapterus leucas from Svalbard during capture.  Journal of the Acoustic Society of America 113: 57-60.
  147. Sørmo, E.G., Skaare, J.U., Lydersen, C., Kovacs, K.M., Hammill, M.O. and Jenssen, B.M.  2003.  Partitioning of persistent organic pollutants in grey seal (Halichoerus grypus) mother-pup pairs.  The Science of the Total Environment 302(1-3): 145-155.
  148. Lydersen, C., Kovacs, K.M., Ries, S. and Knauth, M.  2002.  Precocial diving and patent foramen ovale in bearded seal (Erignathus barbatus) pups. Journal of Comparative Physiology B 172: 713-717.
  149. Lydersen, C., Nøst, O.A., Lovell, P., McConnell, B.J., Gammelsrød, T., Hunter, C., Fedak, M.A. and Kovacs, K.M.  2002.  Salinity and temperature structure of a freezing Arctic fjord – monitored by white whales (Delphinapterus leucas).  Geophysical Research Letters, Vol. 29: No. 23, 2119, doi:10.2029/2002GL015462.
  150. Pomponi, M., Bertonati, C., Patamia, M., Marta, M., Derocher, A.E., Lydersen, C., Kovacs, K.M., Wiig, Ø. and Bårdgard, A.J.  2002.  Polar bear hemoglobin and human HbA: same Binding site but asymmetry of the binding? Hemoglobin 26(4): 363-371.
  151. Hop, H.T. Pearson, Hegseth, E.N., Kovacs, K.M., Wiencke, C., Kwasniewski, S., Eiane, K., Mehlum, F., Gulliksen, B., Wlodarska-Kowalczuk, M., Lydersen, C., Weslawski, J.M., Cochrane, S., Gabrielsen, G.W., Leakey, R.J.G.,  Lønne, O.J., Zajaczkowski, M., Falk-Petersen, S., Kendall, M., Wängberg, S-Å., Bischof, K., Voronkov, A.Y., Kovaltchouk, N.A., Wiktor, J., Poltermann, M., di Prisco, G., Papucci, C. and Gerland, S.  2002.  The marine ecosystem of Kongsfjorden, Svalbard.  Polar Research 21(1): 167-208.
  152. Van Parijs, S.M. and Kovacs, K.M.  2002.  In air and underwater vocalizations of Eastern Canadian harbour seals, Phoca vitulina. Canadian Journal of Zoology 80(7): 1173-1179.
  153. Urashima, T., Sato, H., Munakata, J., Nakamura, T., Arai, I., Saito, T., Tetsuka, M., Fukui, Y., Ishikawa, H., Lydersen, C. and Kovacs, K.M.  2002.  Chemical characterisation of the oligosaccharides in beluga (Dephinapterus leucas) and Minke Whale (Balaenoptera acutorostrata) milk. Comparative Biochemistry and Physiology Part B 132: 611-624.
  154. Karlsen, J.D., Bisther, A., Lydersen, C., Haug, T. and Kovacs, K.M.  2002.  Summer vocalisations of adult male white whales (Delphinapterus leucas) in Svalbard, Norway.  Polar Biology 25: 808-817.
  155. Tittlemier, S., Borrell, A., Duffe, J., Duignan, P.J., Hall, A., Hoekstra, P., Kovacs, K.M., Krahn, M., Lebeuf, M., Lydersen, C., McFee, W., Muir, D., O’Hara, T., Olsson, M., Pranschke, J., Ross, P., Stern, G., Tanabe, S. and Norstrom, R.  2002.  Global distribution of halogenated Dimethyl bipyrroles in marine mammals – evidence of a natural origin. Archives of Environmental Contamination 43(2): 244-255.
  156. Furgal, C., Innes, S. and Kovacs, K.M.  2002.  Inuit spring hunting techniques and local knowledge of the ringed seal in Arctic Bay (Ikpiarjuk), Nunavut.  Polar Research 21(1): 1-16.
  157. Krafft, B.A., Lydersen, C., Gjertz, I. and Kovacs, K.M.  2002.  Diving behaviour of sub-adult harbour seals (Phoca vitulina) at Prins Karls Forland, Svalbard. Polar Biology 25: 230-234.
  158. Lesage, V., Hammill, M.O. and Kovacs, K.M.  2002.  Diet-tissue fractionation of stable carbon and nitrogen isotopes in phocid seals.  Marine Mammal Science 18(1): 182-193.
  159. Jørgensen, C., Lydersen, C., Brix, O. and Kovacs, K.M.  2001.  Diving development in nursing harbour seal pups.  Journal of Experimental Biology 204: 3993-4004.
  160. Van Parijs, S.M., Kovacs, K.M. and Lydersen, C.  2001. Spatial and temporal distribution of vocalizing male bearded seals – implications for male mating strategies. Behaviour 138(7): 905-922.
  161. Lydersen, C., Martin, A.R., Kovacs, K.M. and Gjertz, I.  2001. Summer and autumn movements of white whales (Delphinapterus leucas) in Svalbard, Norway.  Marine Ecology Progress Series 219: 265-274.
  162. Urashima, T., Arita, M., Yoshida, M., Nakamura, T., Aria, I., Saito, T., Arnould, J.P.Y., Kovacs, K.M. and Lydersen, C.  2001.  Chemical characterisation of the oligosaccharides in hooded seal (Cystophora cristata) and Australian fur seal (Arctocephalus pusillus doriferus) milk.  Comparative Biochemistry and Physiology B – Biochemistry & Molecular Biology 128(2): 307-323.
  163. Lesage, V., Hammill, M.O. and Kovacs, K.M.  2001.  Marine mammals and the community structure of the Gulf and Estuary regions of the St. Lawrence (Canada).  Marine Ecology Progress Series 210: 203-221.
  164. Schreer, J.F., Kovacs, K.M. and O’Hara Hines, R.J.  2001.  Comparative diving patterns in pinnipeds and seabirds. Ecological Monographs 71(1): 137-162.
  165. Thompson, P.M., Van Parijs, S. and Kovacs, K.M.  2001.  Local declines in the abundance of harbour seals; implications for the designation and monitoring of protected areas.  Journal of Applied Ecology 38: 117-125.
  166. Lydersen, C., Kovacs, K.M. and Lydersen, E.  2001.  Rust-colored bearded (Erignathus barbatus) and ringed (Phoca hispida) seals from Svalbard, Norway.  Journal of Mammogy 82 (1): 225-230.
  167. Andersen, G., Kovacs, K.M., Lydersen, C., Skaare, J.U., Gjertz, I. and Jenssen, B.M.  2001.  Concentrations and patterns of organochlorine contaminants in white whales (Delphinapterus leucas) from Svalbard, Norway.  Science of the Total Environment 264: 267-281.
  168. Krafft, B.A., Lydersen, C., Kovacs, K.M., Gjertz, I. and Haug, T. 2000.  Diving behaviour of lactating bearded seals (Erignathus barbatus) in the Svalbard area.  Canadian Journal of Zoology 78: 1408-1418.
  169. Gjertz, I., Kovacs, K.M., Lydersen, C. and Wiig, Ø.  2000.  Movements and diving of adult ringed seals (Phoca hispida) in Svalbard.  Polar Biology 23(9): 651-656.
  170. Gjertz, I., Kovacs, K.M., Lydersen, C. and Wiig, Ø.  2000.  Movements and diving of bearded seal (Erignathus barbatus) mother and pups during lactation and post-weaning.  Polar Biology 23(8): 559-566.
  171. Dahl, T.M., Lydersen, C., Kovacs, K.M., Falk-Petersen, S., Sargent, J., Gjertz, I. and Gulliksen, B.  2000.  Fatty acid composition of the blubber in white whales (Delphinapterus leucas).  Polar Biology 23: 401-409.
  172. Lydersen, C., Kovacs, K.M. and Hammill, M.O.  2000.  Reversed molting pattern in starveling gray (Halichoerus grypus) and harp (Phoca groenlandica) seal pups.  Marine Mammal Science 16(2): 489-493.
  173. Bernt, K.E., Hammill, M.O., Lebeuf, M. and Kovacs, K.M.  1999.  Levels and patterns of PCBs and OC pesticides in harbour and grey seals from the St Lawrence Estuary, Canada.  Science of the Total Environment 243-244: 243-262.
  174. Lydersen, C. and Kovacs, K.M.  1999.  Behaviour and energetics of ice-breeding, North Atlantic phocid seals during the lactation period. Marine Ecology Progress Series 187: 265-281.
  175. Debier, C., Kovacs, K.M., Lydersen, C., Mignolet, E. and Larondelle, Y.  1999.  Vitamin E, vitamin A and fatty acid composition of harp and hooded seal milk through lactation.  Canadian Journal of Zoology 77: 952-958.
  176. Lesage, V., Hammill, M.O. and Kovacs, K.M.  1999.  Functional classification of harbour seal (Phoca vitulina) dives using depth profiles, swimming velocity and an index of foraging success.  Canadian Journal of Zoology 77: 74-87.
  177. Marta, M., Patamia, M., Colella, A., Sacchi, S., Pomponi, M., Kovacs, K.M., Lydersen, C. and Giardina, B.  1998.  Anionic binding sites and 2,3-DPG effect in bovine hemoglobin.  Biochemistry 37: 14024-14029.
  178. Schenk, A., Obbard, M.E. and Kovacs, K.M.  1998.  Genetic relatedness and home range overlap among female black bears (Ursus americanus).  Canadian Journal of Zoology 76(8): 1511-1519.
  179. Schreer, J.F., O’Hara Hines, R.J. and Kovacs, K.M.  1998.  Classification of multivariate diving data from air-breathing vertebrates: a comparison of traditional statistical clustering and unsupervised neural networks.  Journal of Agricultural, Biological & Environmental Statistics 3(4): 383-404.
  180. Kovacs, K.M., Lydersen, C., Hammill, M.O., White, B., Wilson, P. and Malik, S.  1997.  A harp seal X hooded seal hybrid.  Marine Mammal Science 13(3): 460-468.
  181. Hammill, M.O., Lydersen, C., Kovacs, K.M. and Sjare, B.  1997.  Estimated fish consumption by hooded seals (Cystophora cristata), in the Gulf of St Lawrence. Journal of Northwest Atlantic Fisheries Science 22: 249-257.
  182. Schreer, J.F. and Kovacs, K.M.  1997.  Allometry of diving behaviour in air-breathing vertebrates.  Canadian Journal of Zoology 75: 339-358.
  183. Lydersen, C., Kovacs, K.M. and Hammill, M.O.  1997.  Energetics during nursing and early postweaning fasting in hooded seal (Cystophora cristata) pups from the Gulf of St Lawrence, Canada.  Journal of Comparative Physiology B 167: 81-88.
  184. Duignan, P.J., Nielsen, O., House, C., Kovacs, K.M., Dufy, N., Early, G., Sadove, S., St. Aubin, D.J., Rima, B.K. and Geraci, J.R.  1997.  Epizootiology of morbillivirus infection in harp seals (Phoca groenlandica), hooded seals (Cystophora cristata), and ringed seals (Phoca hispida) from the Canadian Arctic and Western Atlantic.  Journal of Wildlife Diseases 33(1): 7-19.
  185. Kovacs, K.M., C. Lydersen and M.O. Hammill.  1996.  Grey seal cannibalism.  Mar. Mamm. Sci. 12(1): 161.
  186. Boskovic, R., Kovacs, K.M., Hammill, M.O. and White, B.  1996.  Geographic distribution of mitochondrial DNA haplotypes in grey seals (Halichoerus grypus). Canadian Journal of Zoology 74: 1787-1796.
  187. Lydersen, C., Kovacs, K.M.,  Hammill, M.O. and Gjertz, I.  1996.  Energy intake and utilisation by nursing bearded seal (Erignathus barbatus) pups in Svalbard, Norway.  Journal of Comparative Physiology 166: 405-411.
  188. Lydersen, C. and Kovacs, K.M.  1996.  Energetics of lactation in harp seals (Phoca groenlandica) from the Gulf of St Lawrence, Canada.  Journal of Comparative Physiology 166: 295-304.
  189. Kovacs, K.M., Lydersen, C. and Gjertz, I.  1996.  Birth site characteristics and prenatal molting in bearded seals (Erignathus barbatus).   Journal of Mammalogy 77 (4): 1085-1091.
  190. Haller, M.A., Kovacs, K.M. and Hammill, M.O.  1996.  Maternal behaviour and energy investment by grey seals (Halichoerus grypus) breeding on land-fast ice.  Canadian Journal of Zoology 74: 1531-1541.
  191. Kovacs, K.M., Lydersen, C., Hammill, M.O. and Lavigne, D.M.  1996.  Reproductive effort of male hooded seals (Cystophora cristata): Estimates from mass loss.  Canadian Journal of Zoology 74: 1521-1530.
  192. Furgal, C.M., Kovacs, K.M. and Innes, S.   1996.  Characteristics of ringed seal (Phoca hispida) subnivian lairs and habitat and their effects on predation. Canadian Journal of Zoology 74: 858-874.
  193. Bernt, K.E., Hammill, M.O. and Kovacs, K.M.  1996.  Age estimation in grey seals (Halichoerus grypus) using incisors.  Marine Mammal Science 12: 476-482.
  194. Schenk, A. and Kovacs, K.M.  1996.  Genetic variation in a natural population of black bears (Ursus americanus); revealed by DNA fingerprinting.  Journal of Mammalogy 77(4): 942-950.
  195. Duignan, P.J., Saliki, J.T., St. Aubin, D.J., Early, G., Sadove, S., House, J.A., Kovacs, K.M. and Geraci, J.R.  1995.  Epizootiology of morbillivirus infection in North American Harbor seals (Phoca vitulina) and gray seals (Halichoerus grypus).  Journal of Wildlife Diseases 31(4): 491-501.
  196. Schenk, A. and Kovacs, K.M.  1995.  Multiple mating between black bears revealed by DNA fingerprinting.  Animal Behaviour 50:1483-1490.
  197. Lydersen, C. and Kovacs, K.M.  1995.  Paralysis as a defence response to threatening stimuli in harp seals (Phoca groenlandica).  Canadian Journal of Zoology 73: 486-492.
  198. Kovacs, K.M.  1995.  Mother-pup reunions in harp seals, Phoca groenlandica: cues for the relocation of pups.  Canadian Journal of Zoology 73: 843-849.
  199. Ballard, K.A. and Kovacs, K.M.  1995.  The acoustic repertoire of the hooded seal (Cystophora cristata).  Canadian Journal of Zoology 73: 1362-1374.
  200. Tinker, T.M., Kovacs, K.M. and Hammill, M.O.  1995.  The reproductive behaviour and energetics of male grey seals (Halichoerus grypus) breeding on a landfast ice substrate.  Behavioural Ecology and Sociobiology 36: 159-170.
  201. Lydersen, C., Hammill, M.O. and Kovacs, K.M.  1995.  Milk intake, growth and energy consumption in pups of ice-breeding grey seals (Halichoerus grypus) from the Gulf of St. Lawrence, Canada.  Journal of Comparative Physiology B 164: 585-592.
  202. Thompson, P.M., Kovacs, K.M. and McConnell, B.J.  1994.  Natal dispersal of harbour seals (Phoca vitulina) from breeding sites in Orkney, Scotland.  Journal of Zoology, London 234: 668-673.
  203. Hammill, M.O., Kovacs, K.M. and Lydersen, C.  1994.  Local movements by nursing bearded seal (Erignathus barbatus) pups.  Polar Biology 14:  569-570.
  204. Lydersen, C., Hammill, M.O. and Kovacs, K.M.  1994.  Activity of lactating ice-breeding grey seals (Halichoerus grypus) from the Gulf of St Lawrence, Canada.  Animal Behaviour 48: 1417-1425.
  205. Lydersen, C., Hammill, M.O. and Kovacs, K.M.  1994.  Diving activity in nursing bearded seal (Erignathus barbatus) pups.  Canadian Journal of Zoology 72: 96-103.
  206. McRae, S.B. and Kovacs, K.M.  1994.  Paternity exclusion by DNA fingerprinting and mate guarding in the hooded seal.  Molecular Ecology 3: 101-107.
  207. Pardue, M.T., Sivak, J.G. and Kovacs, K.M.  1993.  Corneal anatomy of marine mammals.  Canadian Journal of Zoology 71: 2282-2290.
  208. Bédard, C., Kovacs, K.M. and Hammill, M.O.  1993.  Cannibalism by grey seals (Halichoerus grypus) on Amet Island, Nova Scotia.  Marine Mammal Science 9(4): 421-424.
  209. Lydersen, C. and Kovacs, K.M. 1993.  Diving behaviour of lactating harp seals (Phoca groenlandica) from the Gulf of St. Lawrence, Canada.  Animal Behaviour 46: 1213-1221.
  210. Kovacs, K.M. and Lavigne, D.M.  1992.  Maternal investment in otariid seals and walruses.  Canadian Journal of Zoology 70: 1953-1964.
  211. Kovacs, K.M. and Lavigne, D.M.  1992.  Mass transfer efficiency between hooded seal (Cystophora cristata) mothers and their pups in the Gulf of St. Lawrence.  Canadian Journal of Zoology 70: 1315-1320.
  212. West, J., Sivak, J.G., Murphy, C.I. and Kovacs, K.M.  1991.  A comparative study of the anatomy of the ciliary body in aquatic mammals.  Canadian Journal of Zoology 69: 2594-2607.
  213. McRae, S.B. and Kovacs, K.M.  1991.  Male specific markers in hooded seal DNA fingerprints. Fingerprint News 3(2): 10-12.
  214. Kovacs, K.M., Lavigne, D.M. and Innes, S.I.  1991.  Mass transfer efficiency between harp seal Phoca groenlandica mothers and their pups during lactation.  Journal of Zoology,  London 223: 213-221.
  215. Kovacs, K.M.  1990.  Mating strategies in male hooded seals (Cystophora cristata).  Canadian Journal of Zoology 68: 2499-2502.
  216. Kovacs, K.M., Jonas, K. and Welke, S.  1990.  Sex and age segregation by Phoca vitulina concolor at haul-out sites during the breeding season in the Passamaquoddy Bay Region, New Brunswick. Marine Mammal Science 6(3): 204-214.
  217. Kovacs, K.M. and Innes, S.I.  1990.  The impact of tourism on harp seals (Phoca groenlandica) in the Gulf of St. Lawrence, Canada.  Applied Animal Behaviour Science 26: 15-26.
  218. Kovacs, K.M.  1987.  Maternal behaviour and early behavioural ontogeny of grey seals (Halichoerus grypus) on the Isle of May, U.K.  Journal of  Zoology, London 213: 697‑715.
  219. Kovacs, K.M.  1987.  Maternal behaviour and early behavioural ontogeny of harp seals (Phoca groenlandica).  Animal Behaviour 35: 844‑855.
  220. Innes, S., Lavigne, D.M., Earle, W.M. and Kovacs, K.M.  1987.  Feeding rates of seals and whales.  Journal of Animal Ecology 56: 115‑130.
  221. Lavigne, D.M., Innes, S., Worthy, G.A.J. and Kovacs, K.M.  1986.  Metabolic rate ‑ body size relations in marine mammals.  Journal of Theoretical Biology 122: 123‑124.
  222. Kovacs, K.M. and Lavigne, D.M.  1986.  Growth of grey seal (Halichoerus grypus) neonates: differential maternal investment in the sexes.  Canadian Journal of Zoology 64: 1937‑1943.
  223. Kovacs, K.M. and Lavigne, D.M.  1986.  Maternal investment and neonatal growth in phocid seals.  Journal of Animal Ecology 55: 1035‑1051.
  224. Innes, S., Lavigne, D.M., Earle, W.M. and Kovacs, K.M.  1986.  Estimating feeding rates of marine mammals from heart mass to body mass ratios.  Marine Mammal Science 2(3): 227‑229.
  225. Lavigne, D.M., Innes, S., Worthy, G.A.J., Kovacs, K.M., Schmitz, O.J. and Hickie, J.  1986.  Metabolic rates of seals and whales.  Canadian Journal of Zoology 64: 279‑284.
  226. Kovacs, K.M., Lavigne, D.M. and Stewart, R.E.A.  1985.  Early postnatal mortality in Northwest Atlantic harp seals (Phoca groenlandica).  Journal of Mammalogy 66: 556‑558.
  227. Kovacs, K.M. and Lavigne, D.M.  1985.  Neonatal growth and organ allometry of Northwest Atlantic harp seals (Phoca groenlandica).  Canadian Journal of Zoology 63: 2793‑2799.
  228. Kovacs, K.M. and Ryder, J.P.  1985.  Morphology and physiology of female­-female pair members.  Auk 102: 874-878.
  229. Kovacs, K.M. and Ryder, J.P.  1983.  Reproductive performance of female-female pairs and polygynous trios of Ring‑billed Gulls.  Auk 100: 658‑669.
  230. Kovacs, K.M. and Ryder, J.P.  1981.  Nest site‑tenacity and mate fidelity in female‑female pairs of Ring‑billed Gulls.  Auk 93: 625‑627.